Gnaphosa zonsteini, Fomichev, Alexander A. & Marusik, Yuri M., 2017

Fomichev, Alexander A. & Marusik, Yuri M., 2017, A survey of East Palaearctic Gnaphosidae (Araneae). 8. New data on Berlandina and Gnaphosa from Mongolia, Zootaxa 4258 (1), pp. 69-80 : 75-77

Gnaphosa zonsteini sp. n.

Figs 20‒27 , 32‒33 , 35

Material. MONGOLIA: Khovd Aimag: holotype ♂ ( ISEA) and paratypes 4♂ 1♀ ( ISEA) 36 km SW from Altai Village, Bodonchiyn-Gol River Valley , 45°46’N, 92°12’E, tugay (gallery forest), night, attracted by light, 1280 m, 17‒ 18.05.2015 (AF) ; 1♂ ( ZMMU) same locality, 1.06.2015 (AF).

Etymology. The species name is a patronym in honour of our friend and colleague Sergei L. Zonstein, on the occasion of his 60th birthday, the 8th of August, 2016.

Diagnosis. Males of the new species resemble G. esyunini Marusik, Fomichev & Omelko, 2014 by having large embolic spine directed prolaterally, but can be distinguished from the former species by longer embolus without serration on the prolateral edge, lowered base of embolus and not bulged median apophysis (cf. Figs 21–27 and figs 7–11 in Marusik et al. 2014b). Females similar to G. gracilior Kulczyński, 1901 , G. muscorum (L. Koch, 1866) , G. rasnitsyni Marusik, 1993 and G. wiehlei Schenkel, 1963 , but differ from the later three species by the shape of atrium and septum and from G. gracilior by less concave posterior margin of scape and head of receptacles directed upward.

Description. Male. Total length 6.6. Carapace: 3.0 long, 2.3 wide. Coloration. Prosoma and chelicerae darkbrown. Carapace with black edges and two dark stripes around cephalic area. Legs and palps brown. Abdomen dark-gray, almost black dorsally and yellow-gray ventrally. Spinnerets gray.

Leg measurements:

Fe Pt Ti Mt Ta Total I 2.1 1.2 1.65 1.5 1.25 7.7 II 1.9 1.1 1.45 1.35 1.2 7.0 III 1.75 0.95 1.2 1.5 1.05 6.45 IV 2.35 1.2 1.85 2.5 1.35 9.25 Leg spination:

Fe Pt Ti Mt

I d1-1-0 p0-0- 1 v 1-1- 2 v 2-0-0

II d1-1-0 p0-1-1 p0-0- 1 v 1-2- 2 v 2-0-0

III d1-1-0 p0-1-1 r0-1-1 r1-0 d1-0-0 p2-1-1 r2-1- 1 v 2-2-2 d0-1-0 p1-1-0 r1-1-0 v2 -0- 2 IV d1-1-0 p0-1-1 r0-1-1 p2-1-1 r2-1- 1 v 2-2-2 d0-2-0 p1-1-0 r1-1-0 v2-2 -0 Palp as in Figs 21‒27 ; tibial apophysis rather long, almost equal in length to tibia, with pointed tip; median apophysis large, hook-shaped; embolus long, filamentous, embolic base with strong spine directed prolaterally. Size variations. Total length 6.2–8.3; carapace 3.0–3.65 long and 2.3–2.9 wide (n=5). Female. Total length 12.3. Carapace: 4.35 long, 3.25 wide. Coloration as in male, but with lighter carapace and monochrome dirty-yellow abdomen.

Leg measurements:

Fe Pt Ti Mt Ta Total I 2.65 1.6 1.95 1.55 1.3 9.05 II 2.4 1.45 1.7 1.45 1.2 8.2

III 2.2 1.25 1.4 1.8 1.15 7.8

IV 2.95 1.6 2.15 2.85 1.4 10.95

Leg spination:

Fe Pt Ti Mt

I d1-0-0 p0-0- 1 v 0-0- 2 v 0-2-0

II d1-1-0 p0-0- 1 v 0-1- 2 v 0-2-0

III d0-1-0 p0-1-1 r0-1-1 r1-0 p2-1-1 r1-1- 1 v 2-2-2 d0-1-0 p1-1-0 r1-1-0 v2-1 -0 IV d1-1-0 p0-0-1 r0-0-1 p1-1-1 r2-1- 1 v 2-2-2 d0-1-0 p1-1-0 r1-0-0 v2-2 -0 Epigyne as in Figs 32‒33 ; fovea diamond-shaped, with pair of clearly-delineated bulges (Ab); scape wider than long with slightly concave posterior margin and stretched pocket of scape (Ps); head of receptacles (Rh) directed upwards.

Distribution. Known from the type locality only.

Comments. Judging from the combination of such features as base of embolus with spine in anterior part, filiform and rounded embolus and hook-like median apophysis it can be assumed that G. zonsteini sp. n. belongs to a monophyletic species group formed by following species: G. esyunini Marusik, Fomichev & Omelko, 2014 , G. khovdensis Marusik, Fomichev & Omelko, 2014 and G. rasnitsyni Marusik, 1993 (see below).

Figures

FIGURES 20 - 27. Male of Gnaphosa zonsteini sp. n. 20 habitus, dorsal; 21 ‒ 23 palp, ventral, retro- and prolateral; 24 cymbium and tibia, retrolateral; 25 ‒ 27 bulb, prolateral, ventral and anterior. Scale = 0.2 mm if not otherwise indicated.

FIGURES 28 ‒ 33. Female of Gnaphosa rasnitsyni (28 ‒ 31) and G. zonsteini sp. n. (32 ‒ 33). 28 habitus, dorsal; 29 intact epigyne, ventral; 30, 32 macerated epigyne, ventral; 31, 33 macerated epigyne, dorsal. Scale = 0.2 mm if not otherwise indicated. Abbreviations. Ab foveal bulges, Lp lateral pockets, Ps pocket of the scape; Rh head of receptacle.

FIGURES 34 ‒ 35. Collecting localities of Berlandina and Gnaphosa species examined in this study. Solid symbol on Fig. 34 refers to new records, open symbols to the literature data, dotted square refers to the region shown on Fig. 35. Blue squares in Fig. 35 refer to the records of G. ustyuzhanini (= G. rasnitsyni). One symbol may designate more than one locality.