Limonium species

Brullo, Salvatore & Erben, Matthias, 2016, The genus Limonium (Plumbaginaceae) in Greece, Phytotaxa 240 (1), pp. 448-450 : 448-450

Key to the Limonium species occurring in Greece

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- Leaves sinuate; stems winged; calyx blue ............................................................................................................................. 2 Leaves entire; stems not winged; calyx white........................................................................................................................ 3 Plant annual; calyx seam with very large lobes .................................................................................................. 2. L. lobatum Plant perennial; calyx seam nearly undivided................................................................................................... 1. L. sinuatum Plant annual; calyx teeth very long and narrow, with decurrent nerves ............................................................ 3. L. echioides Plant perennial; calyx teeth short with no decurrent nerves .................................................................................................. 4 Leaves very large, 5–40 × 3–9 cm, usually pinnately nerves ................................................................................................ 5 Leaves small, 0.5–10 × 0.1–2.5 cm, not pinnately nerves ..................................................................................................... 9 Inner bract 2.6–3.7 mm long, seldom until 3.8 mm long, than leaves dotted on both .......................................................... 6 Inner bract 3.8–5.8 mm long .................................................................................................................................................. 7 Spikes with 7–12 spikelets per cm; spikelets 1-flower.................................................................................. 7. L. compactum Spikes with 4–6 spikelets per cm; spikelets 2–3-flowers .............................................................................. 9. L. hirsuticalyx Inner bract 4.8–5.8 mm long .......................................................................................................................... 8. L. pagasaeum Inner bract 3.7–4.5 mm long .................................................................................................................................................. 8 Spikelets 4.5–6.0 mm long; calyx 3.9–5.0 mm long...................................................................................... 6. L. narbonense Spikelets 6.2–9.0 mm long; calyx 5.2–5.8 mm long ............................................................................. 10. L. brevipetiolatum Inner bract hairy ................................................................................................................................................................... 10 Inner bract glabrous.............................................................................................................................................................. 14 Inner bract totally hairy. ............................................................................................................................. 13. L. ammophilon Inner bract only in the upper part hairy.................................................................................................................................11 Spikelets 5.5–7.0 mm long; calyx 4.5–5.2 mm long...................................................................................... 44. L. cythereum Spikelets 7.5–9.5 mm long; calyx 5.8–7.0 mm long............................................................................................................ 12 Leaves spathulate; outer and inner bracts with crateriform glands ................................................................... 14. L. roridum Leaves oblanceolate; outer and inner bracts rugose............................................................................................................. 13 Outer bract hairy; spikelets 8.5–9.5 mm long................................................................................................ 16. L. dolihiense Outer bract glabrous; spikelets 7.5–8.5 mm long...................................................................................... 22. L. oligotrichum

LIMONIUM (PLUMBAGINACEAE)

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- Stems hairy ........................................................................................................................................................................... 15 Stems glabrous ..................................................................................................................................................................... 17 Plant densely pubescent ........................................................................................................................ 29. L. damboldtianum Only stems hairy near the base............................................................................................................................................. 16 Sterile branches numerous; inner bracts 3.6–4.1 × 2.5–3.0 mm ......................................................................... 39. L. pylium Sterile branches few; inner bracts 4.2–5.0 × 3.2–3.5 mm ........................................................................ 28. L. cephalonicum Stems tuberculate with many raised crateriform glands (Fig. II, type A) ............................................................................ 18 Stems verrucated with crescent-shaped glands (Fig. II, type B), rugose with deepened glands (Fig. II, type C) or smooth .. .............................................................................................................................................................................................. 53 Spikelets 4.5–8.5 mm long................................................................................................................................................... 19 Spikelets 8.5–11.5 mm long ................................................................................................................................................. 39 Inner bract 3.8–5.2 mm long ................................................................................................................................................ 20 Inner bract 5.7–8.0 mm long ................................................................................................................................................ 21 Sterile branches absent or few; inner bract 3.2–3.9 mm broad ................................................................. 42. L. cornarianum Sterile branches numerous; inner bract 2.6–2.9 mm broad .......................................................................... 36. L. zacynthium Plant 3–8 cm tall ............................................................................................................................................... 50. L. pusillum Plant more than 10 cm tall ................................................................................................................................................... 22 Leaves linear, 5–14 cm long, semi-cylindrical in cross-section .................................................................... 20. L. carpathum Leaves narrowly oblanceolate to spathulate, normally shorter than 4.5 cm, flat or V-shaped in cross-section................... 23 Spikelets 6.5–7.2 mm long................................................................................................................................................... 24 Spikelets 7.2–9.0 mm long................................................................................................................................................... 25 Inner bract 6.4–6.7 × 3.9–4.4 mm ..................................................................................................................... 49. L. palmare Inner bract 5.7–6.3 × 3.4–3.9 mm .................................................................................................................. 52. L. parosicum Leaves 1–5 mm broad .......................................................................................................................................................... 26 Leaves 5–12 mm broad ........................................................................................................................................................ 33 Inner bract 7.3–8.0 mm long ................................................................................................................................................ 27 Inner bract 5.8–7.2 mm long ................................................................................................................................................ 28 Calyx 5.0– 6.1 mm long ...........................................................................................................................54. L. elaphonisicum Calyx 6.1–7.0 mm long ...................................................................................................................................... 56. L. xiliense Inner bract 6.6–7.2 mm long ................................................................................................................................................ 29 Inner bract 5.8–6.5 mm long ................................................................................................................................................ 32 Sterile branches numerous ........................................................................................................................ 53. L. chersonesum Sterile branches absent or few.............................................................................................................................................. 30 Spikelets 7.5–8.0 mm long .................................................................................................................. 18. L. xerocamposicum Spikelets 8.1–9.0 mm long................................................................................................................................................... 31 Outer bract 2.0– 2.3 mm long; inner bract 2.8–3.0 mm broad............................................................................ 60. L. sougiae Outer bract 2.6–3.2 mm long; inner bract 3.6–4.1 mm broad ............................................................................ 56. L. xiliense Stems 5–10 cm long; few sterile branches; inner bract 5.8–6.1 mm long ....................................................... 55. L. ikaricum Stems 15–45 cm long; several sterile branches; inner bract 6.1–6.5 mm long .................................................... 57. L.quinnii Inner bract 4.9–5.2 mm broad..................................................................................................................... 70. L. vravronense Inner bract 3.1–4.8 mm broad.............................................................................................................................................. 34 Sterile branches absent or few (1–3 per stem)................................................................................................... 27. L. samium Sterile branches numerous ................................................................................................................................................... 35 Inner bract 7.3–8.2 mm long ................................................................................................................................................ 36 Inner bract 6.0– 7.2 mm long ................................................................................................................................................ 38 Leaves fleshy, green, verrucated .............................................................................................................54. L. elaphonisicum Leaves coriaceous, glaucous, with many crateriform glands............................................................................................... 37 Living leaves in terminal rosettes, V-shaped in cross section ....................................................................... 62. L. proliferum Living leaves distributed in the upper two thirds, flat in cross-section................................................................ 64. L. thirae Leaves fleshy; inner bract verrucated; ribs of the calyx ending at the base of the lobes ................................... 69. L. atticum Leaves coriaceous; inner bract with many crateriform glands; ribs of the calyx ending in the middle of the lobes............... ....................................................................................................................................................................... 62. L. proliferum Inner bract 6.3–7.2 mm long ................................................................................................................................................ 40 Inner bract 7.2–10.5 mm long .............................................................................................................................................. 42 Leaves small oblanceolate, apex acute to obtuse; inner bract 2.8–3.0 mm broad............................................. 60. L. sougiae Leaves oblong-spathulate to spathulate, apex obtuse to rounded or emarginate; inner bract 3.2–4.5 mm broad................ 41 Leaves fleshy; inner bract verrucated; ribs of calyx ending at the base of lobes .............................................. 69. L. atticum Leaves coriaceous; inner bract with many crateriform glands; ribs of calyx ending in the middle of lobes........................... ....................................................................................................................................................................... 62. L. proliferum Sterile branches absent or few (1–4 per stem) ..................................................................................................................... 43 Sterile branches numerous (rarely few) ............................................................................................................................... 47

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71 Inner bract 8.4–9.5 mm long............................................................................................................................. 21. L. sitiacum Inner bract 7.1–8.2 mm long ................................................................................................................................................ 44 Leaves smooth, dotted; axes of spikes without crateriform glands.................................................................... 67. L. taenari Leaves and axes of spikes with many crateriform glands.................................................................................................... 45 Leaves linear-oblanceolate (L/B = 10–15) ......................................................................................................... 56. L. xiliense Leaves narrowly oblanceolate to spathulate (L/B = 5–8.5) ................................................................................................. 46 Stems numerous; inflorescence narrowly trullate to narrowly obtrullate; bracts with crateriform glands .... 65. L. recticaule Stems few; inflorescence broadly obtriangular to obtrullate; bracts without crateriform glands.................. 23. L. amopicum Inner bract 8.2–9.0 mm long ................................................................................................................................................ 48 Inner bract 7.0–8.2(–8.4) mm long ...................................................................................................................................... 49 Calyx 6.0– 7.2 mm long .............................................................................................................................. 6 6. L. crateriforme Calyx 7.8–8.3 mm long ................................................................................................................................ 68. L. schinousae Leaves and stems green; leaves without crateriform glands ...................................................................54. L. elaphonisicum Leaves and stems glaucous; leaves with crateriform glands................................................................................................ 50 Leaves 5.0– 10 mm broad, spathulate to oblanceolate-spathulate ........................................................................................ 51 Leaves 1.5–5.0 mm broad, linear to narrowly oblanceolate or narrowly spathulate ........................................................... 52 Living leaves in terminal rosettes, V-shaped in cross-section ....................................................................... 62. L. proliferum Living leaves distributed in the upper two thirds of caudices, flat in cross-section............................................. 64. L. thirae Stems several to numerous; leaves fleshy, linear to narrowly oblanceolate; sterile branches several to numerous ................ .......................................................................................................................................................................63. L. stenotatum Stems few; leaves coriaceous, narrowly spathulate; sterile branches absent or only 1–6 per stem............... 23. L. amopicum Sterile branches several (at least 7 per stem) to numerous .................................................................................................. 54 Sterile branches absent or few (1–6 per stem) ..................................................................................................................... 79 Branches reflexed; branching angles 100°–165° ................................................................................................................. 55 Branches obliquely directed upwards; branching angles 30°–90°....................................................................................... 56 Leaves 2–7 mm broad; spikelets 6.5–7.5 mm long.................................................................................... 31. L. antipaxorum Leaves 6-16 mm broad; spikelets 5.0– 6.5 mm long ....................................................................................... 32. L. arcuatum Spikelets 4.9–5.5 mm long................................................................................................................................................... 57 Spikelets 5.9–9.5 mm long................................................................................................................................................... 58 Inner bract 3.2–3.8 × 2.3–2.5 mm .............................................................................................................38. L. messeniacum Inner bract 4.2–5.0 × 3.2–3.5 mm ............................................................................................................ 28. L. cephalonicum Inner bract 6.1–8.0 mm. long ............................................................................................................................................... 59 Inner bract 2.5–6.1 mm long ................................................................................................................................................ 67 Ribs of the calyx ending above the base of the lobes .......................................................................................................... 60 Ribs of the calyx ending before or at the base of the lobes.................................................................................................. 63 Leaves 4–8 mm broad, spathulate to oblanceolate, obtuse to rounded at the apex.............................................................. 61 Leaves 1–4 mm broad, narrowly oblanceolate to narrowly spathulate, acute to obtuse at the apex ................................... 62 Inner bract 3.8–4.2 mm broad, with crateriform glands; spikelets straight .................................................. 62. L. proliferum Inner bract 4.2–4.8 mm broad, rugose; spikelets laterally slightly curved .........................................19. L. microcycladicum Spikelets 7.5–8.0 mm long; calyx tube hairy ............................................................................................ 53. L. chersonesum Spikelets 8.2–9.5 mm long; calyx tube glabrous ................................................................................................ 57. L. quinnii Calyx 5.0– 5.9 mm long........................................................................................................................................................ 64 Calyx 5.9–7.5 mm lon.......................................................................................................................................................... 65 Leaves spathulate to oblanceolate, 6–18 mm broad; inner bract 5.2–7.1 mm long ............................................ 40. L. sieberi Leaves narrowly oblanceolate, 2–5 mm broad; inner bract 7.3–8.0 mm long ........................................ 54. L. elaphonisicum Leaves 3–6 mm broad; inner bract 7.5–8.0 mm long....................................................................................... 75. L. helenae Leaves 6–18 mm broad; inner bract 6.1–7.3 mm long ........................................................................................................ 66 Sterile branches absent or few (1–5 per stem); calyx exceeding the inner bract by 1.5–2.5 mm ....................... 40. L. sieberi Sterile branches more than 6 per stem; calyx exceeding the inner bract by 0.5–1.5 mm ................................ 12. L. aegaeum Outer bract totally silver-white; central part of the inner bract without acumen; calyx 2.8–3.8 mm long; inner bract 2.5–3.4 mm long ........................................................................................................................................................4. L. bellidifolium Outer bract white, green or brown; central part of the inner bract with acumen; calyx more 4.4 mm long; inner bract more 3.5 mm long.......................................................................................................................................................................... 68 Inner bract 3.5–4.1 mm long ................................................................................................................................................ 69 Inner bract 4.3–6.1 mm long ................................................................................................................................................ 70 Sterile branches few to several; outer bract 0.9–1.2 mm long; inner bract 2.8–3.2 mm broad ...................... 33. L. ithacense Sterile branches very numerous; outer bract 1.2–1.5 mm long; inner bract 2.3–2.7 mm broad (Peloponnese) ...................... ........................................................................................................................................................................ 34. L. coronense Spikelets 7.5–9.0 mm long................................................................................................................................................... 71 Spikelets 5.8–7.4 mm long................................................................................................................................................... 73 Sterile branches very numerous; inner bract 4.3–5.2 × 3.0– 3.3 mm .......................................................... 30. L. phitosianum

LIMONIUM (PLUMBAGINACEAE)

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99 Sterile branches few or several; inner bract 5.2–6.1 × 3.4–5.2 mm ..................................................................................... 72 Inner bract 3.4–4.3 mm broad; calyx glabrous or sparsely to densely hairy. ...................................................... 40. L. sieberi Inner bract 4.9–5.2 mm broad; calyx densely, long hairy............................................................................ 70. L.vravronense Plant 5–15 cm tall; stems slender, 3–12 cm long ................................................................................................................. 74 Plant 15–60 cm tall; stems more or less robust, 14–50 cm long.......................................................................................... 75 Outer bract 1.2–1.7 mm long; middle bract 1.5–1.9 mm long............................................................................. 15. L. fragile Outer bract 1.9–2.1 mm long; middle bract 1.9–2.1 mm long ........................................................................ 17. L. athinense Leaf margin revolute; inner bract 3.6–4.8 mm long ............................................................................................................ 76 Leaf margin flat; inner bract 4.8–6.1 mm long .................................................................................................................... 77 Stems very slender; inner bract narrowly elliptic, 2.6–2.9 mm broad; axes of spikes straight, with many crateriform glands ...................................................................................................................................................................... 36. L. zacynthium Stems more or less robust; inner bract elliptic to obovate, 3.0– 3.3 mm broad; axes of spikes flexuous, rugose .................... .................................................................................................................................................................... 35. L. saracinatum Spikelets laterally slightly curved, arranged with 4–6 per cm; leaves 4.0– 8.5 mm broad ............................... 11. L. virgatum Spikelets straight, arranged with 2–4 per cm; leaves 2–4 mm broad................................................................................... 78 Sterile branches very numerous; inner bract 4.8–5.1 × 3.1–3.6 mm; calyx 4.4–4.9 mm long ................ 78. L. astypaleanum Sterile branches few to several; inner bract 5.2–5.8 × 2.7–3.1 mm; calyx 4.9–5.8 mm long .................... 79. L. meandrinum Spikelets 3.5–5.3 mm long................................................................................................................................................... 80 Spikelets 5.5–10.0 mm long................................................................................................................................................. 82 Leaves green, margin revolute ...................................................................................................................... 74. L. aphroditae Leaves grey-green, margin flat............................................................................................................................................. 81 Inner bract 2.6–3.7 mm long; leaves awned, on both sides dotted................................................................ 7. L. compactum Inner bract 3.7–4.0 mm long; leaves without an awn, smooth ................................................................. 73. L. corinthiacum Spikelets 8.3–10.0 mm long................................................................................................................................................. 83 Spikelets 5.5–8.2 mm long................................................................................................................................................... 99 Inner bract 5.9–7.2 mm long ................................................................................................................................................ 84 Inner bract 7.3–9.0 mm long ................................................................................................................................................ 88 Inner bract 2.8–3.0 mm broad ............................................................................................................................60. L. sougiae Inner bract 3.2–4.8 mm broad.............................................................................................................................................. 85 Leaves obcordate to broadly spathulate, apex emarginate................................................................................ 81. L. frederici Leaves narrowly spathulate to oblanceolate, apex obtuse to round ..................................................................................... 86 Leaves 2–5 mm broad; plant 7–20 cm tall ....................................................................................................... 58. L. graecum Leaves 6–18 mm broad; plant 20–40 cm tall ....................................................................................................................... 87 Inner bract 3.4–4.3 mm broad; leaves rugose ..................................................................................................... 40. L. sieberi Inner bract 4.5–5.0 mm broad; leaves smooth, dotted ............................................................................... 47. L. sartorianum Stems articulate; calyx exceeding the inner bract by 1.5–2.0 mm...................................................................... 67. L. taenari Stems not articulate; calyx exceeding the inner bract by 0.5–1.0 mm................................................................................. 89 Outer and inner bracts with crateriform glands.................................................................................................. 27. L. samium Outer and inner bracts rugose .............................................................................................................................................. 90 Leaves smooth to slightly rugose ......................................................................................................................................... 91 Leaves verrucated or tuberculate with crateriform glands................................................................................................... 93 Leaves and stems green; outer bract 2.7–3.0 mm broad; inner bract 4.2–4.6 mm broad.................................. 75. L. helenae Leaves and stems glaucous; outer bract 1.8–2.7 mm broad; inner bract 3.7–4.2 mm broad ............................................... 92 Outer bract 1.8–2.1 mm broad; inner bract 7.8–8.3 mm long; ribs of calyx ending near the upper margin of lobes.............. ......................................................................................................................................................................25. L. calliopsium Outer bract 2.2–2.7 mm broad; inner bract 8.5–9.5 mm long; ribs of calyx ending at the base of lobes......... 21. L. sitiacum Calyx 5.0– 6.2 mm long........................................................................................................................................................ 94 Calyx 6.3–7.8 mm long........................................................................................................................................................ 96 Inner bract 8.4–9.5 mm long............................................................................................................................. 21. L. sitiacum Inner bract 6.0–8.0 mm long ................................................................................................................................................ 95 Spikes with 2–4 spikelets per cm; stems 3–13 cm long; ribs of calyx ending above the base of lobes .......... 58. L. graecum Spikes with 0.5–2 spikelets per cm; stems 20–40 cm long; ribs of calyx ending before or at the base of lobes .............................................................................................................................. 5 4. L. elaphonisicum Calyx 7.2–7.8 mm long; axe of spike without crateriform glands ........................................................... 24. L. monolithicum Calyx 5.7–7.2 mm long; axe of spike with crateriform glands ............................................................................................ 97 Inner bract 4.3–4.8 mm broad; calyx tube glabrous or sparsely hairy; calyx ribs ending at the base of lobes........................ .......................................................................................................................................................................63. L. stenotatum Inner bract 3.2–4.3 mm broad; calyx tube sparsely to densely hairy; calyx ribs ending above the base of lobes............... 98 Stems 3–13 cm long; spikes 1–3.5 cm long, with 2–4 spikelets per cm .......................................................... 58. L. graecum Stems 15–35 cm long; spikes 4–10 cm long, with 0.5–2 spikelets per cm ........................................... 59. L. spreitzenhoferi Spikelets 7.4–8.2 mm long................................................................................................................................................. 100

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- Spikelets 5.5–7.3 mm long..................................................................................................................................................111

100 Leaves 1–5 mm broad ........................................................................................................................................................ 101

- Leaves 5–18 mm broad ...................................................................................................................................................... 105

101 Leaves tuberculate with many crateriform glands or verrucated ....................................................................................... 102

- Leaves smooth or rugose.................................................................................................................................................... 104

102 Stems robust, 20–40 cm long; spikes with 0.5–2 spikelets per cm; ribs of calyx ending before or at the base of lobes......... ................................................................................................................................................................. 54. L. elaphonisicum

- Stems slender, 3–13 cm long; spikes with 2–4 spikelets per cm; ribs of calyx ending above the base of lobes ............... 103

103 Calyx 5.0– 5.5 mm long; outer bract 1.8–2.0 mm broad ..................................................................... 18. L. xerocamposicum

- Calyx 5.7–7.1 mm long; outer bract 2.0–3.0 mm broad .................................................................................. 58. L. graecum

104 Leaves oblong-spathulate, 1.5–4 cm long; inner bract 2.8–3.2 mm broad.................................................. 80. L. kirikosicum

- Leaves linear-oblanceolate, 5–14 cm long; inner bract 3.2–4.0 mm broad .................................................. 20. L. carpathum

105 Inner bract 7.1–7.5 mm long ....................................................................................................................... 26. L. vanandense

- Inner bract 5.1–7.0 mm long .............................................................................................................................................. 106

106 Inner bract 5.1–5.9 mm long .............................................................................................................................................. 107

- Inner bract 5.9–7.0 mm long .............................................................................................................................................. 108

107 Leaves 4–7 mm broad; inner bract 3.2–3.8 mm broad.............................................................................. 77. L. archeothirae

- Leaves 9–20 mm broad; inner bract 4–5 mm broad .................................................................................. 71. L. ocymifolium

108 Inner bract 3.3–4.4 mm broad............................................................................................................................................ 109

- Inner bract 4.5–5.0 mm broad.............................................................................................................................................110

109 Leaves verrucated to tuberculate with crateriform glands; spikes with 0.5–2 spikelets per cm; calyx exceeding the inner bract by 1.0– 1.5 mm ........................................................................................................................................ 61. L. grabusae

- Leaves rugose; spikes with 2–3 spikelets per cm; calyx exceeding the inner bract by 1.5–2.5 mm .................. 40. L. sieberi

110 Outer bract 3.0– 3.8 mm long; fertile branches 1–3 cm long; spikes with 3–4 spikelets per cm .................... 85. L. sirinicum

- Outer bract 2.5–3.0 mm long; fertile branches 3–10 cm long; spikes with 0.5–3 spikelets per cm ......... 47. L. sartorianum

111 Spikelets 6.5–7.3 mm long..................................................................................................................................................112

- Spikelets 5.5–6.4 mm long................................................................................................................................................. 124

112 Leaves 8–20 mm broad .......................................................................................................................................................113

- Leaves 2–7 mm broad .........................................................................................................................................................119

113 Leaves acute to obtuse, with margin broadly white............................................................................. 84. L. albomarginatum

- Leaves rounded to obtuse, without margin white...............................................................................................................114

114 Inner bract 6.2–7.1 mm long ...............................................................................................................................................115

- Inner bract 4.8–6.1 mm long ...............................................................................................................................................116

115 Leaves coriaceous; inner bract 4.1–4.8 mm broad; calyx exceeding the inner bract by 0.5–1.5 mm ............. 43. L. creticum

- Leaves fleshy; inner bract 3.4–4.3 mm broad; calyx exceeding the inner bract by 1.5–2.5 mm ........................ 40. L. sieberi

116 Fertile branches 10–30 cm long; spikes with 0.5–2.0 spikelets per cm....................................................... 41. L. hierapetrae

- Fertile branches 8–10 cm long; spikes with 2–5 spikelets per cm......................................................................................117

117 Caudices 6–14 cm long; leaves verrucated; inner bract 5.5–6.0 mm long ...................................................48. L. pigadiense

- Caudices 0.5–6.0 cm long; leaves smooth; inner bract 5.0– 5.5 mm long...........................................................................118

118 Stems smooth; calyx exceeding the inner bract by 1.0– 1.5 mm ................................................................ 71. L. ocymifolium

- Stems rugose to verrucated; calyx exceeding the inner bract by 1.5–2.5 mm .................................................... 40. L. sieberi

119 Leaves linear-oblanceolate, 2–3 mm broad; inner bract 2.7–3.1 mm broad.............................................. 79. L. meandrinum

- Leaves spathulate, (3–) 4–7 mm broad; inner bract 3.2–4.8 mm broad.............................................................................. 120

120 Plant 4–8 cm tall; stems 3–7 cm long ..........................................................................................................51. L. contractum

- Plant 10–50 cm tall; stems 7–40 cm long .......................................................................................................................... 121

121 Stems smooth; inner bract 4.8–5.2 mm long ................................................................................................ 46. L. chrisianum

- Stems rugose or verrucated; inner bract 5.2–6.1 mm long................................................................................................. 122

122 Leaf margin revolute; inner bract 5.2–5.5 mm long.................................................................................. 77. L. archeothirae

- Leaf margin flat; inner bract 5.6–6.3 mm long .................................................................................................................. 123

123 Leaves with many crateriform glands or verrucated with many crescent-shaped glands; outer bract 2.7–3.0 mm long ........ ........................................................................................................................................................................52. L. parosicum

- Leaves only above slightly verrucated; outer bract 2.2–2.6 mm long ..........................................................48. L. pigadiense

124 Spikes with 8–12 spikelets per cm ................................................................................................................ 5. L. glomeratum

- Spikes with 1–7 spikelets per cm ....................................................................................................................................... 125

125 Inner bract 3.5–4.2 mm long .............................................................................................................................................. 126

- Inner bract 4.3–5.3 mm long .............................................................................................................................................. 127

126 Outer bract 0.9–1.2 mm long; inner bract 2.8–3.2 mm broad ......................................................................... 33. L. ithacense

- Outer bract 1.2–1.8.mm long; inner bract 2.0– 2.6 mm broad ....................................................................... 37. L. kardamylii

127 Leaves 2–5 mm broad; inner bract 2.7–2.9 mm broad ................................................................................... 17. L. athinense

- Leaves 5–18 mm broad; inner bract 2.9–4.4 mm broad .................................................................................................... 128

128 Outer bract 1.3–1.7 mm long; inner bract 2.9–3.1 mm broad..................................................................... 82. L. heraionense