Paramblynotus nebulosus, Liu, Ronquist & Nordlander, 2007

Liu, Zhiwei, Ronquist, Fredrik & Nordlander, Göran, 2007, The Cynipoid Genus Paramblynotus: Revision, Phylogeny, And Historical Biogeography (Hymenoptera: Liopteridae), Bulletin of the American Museum of Natural History 2007 (304), pp. 1-151 : 115-117

Paramblynotus nebulosus , new species

FEMALE: Length 5.0–8.0 mm. Body entirely black. Antenna brown and darker toward apex. Legs black, except fore- and middle tibiae and tarsi brown. Wings slightly tinted yellow and with a broad dark gray band along outer margin of forewing. Body pubescence golden-tinted, except pubescence on mesopleural triangle white.

Antenna 13-segmented, flagellum filiform, not widened apically. Vertex foveate-reticulate. Eye extended laterally, distinctly beyond outer margin of gena. Ocellar plate slightly raised and foveate-rugose and defined laterally by superficial carina lined by irregular large foveae along interior side. Median frontal carina briefly present above antennal sockets, raised into a low, rounded process in lateral view. Upper face rugose-foveate/punctate laterally; antennal scrobe distinctly depressed, defined by a thin carina laterally, and coarsely punctate anteriorly and glabrate with longitudinal carination posteriorly. Gena coarsely foveate-reticulate with dense, appressed pubescence; lower face and clypeus coarsely foveate and punctate with dense appressed pubescence; anterior tentorial pits small and distinct; clypeo-pleurostomal sulcus and epistomal sulcus form a superficial, smoothly curved arch. Lateral occipital carina not reaching vertex. Occiput longitudinally carinate medially and glabrous laterally.

Anterior flange of pronotum finely coriarious; submedian pronotal depressions separated from each other medially; anterior pronotal plate glabrous and finely, densely punctate with appressed pubescence. Pronotum raised dorsomedially; pronotal crest gradually raised medially into a peak higher than mesoscutum. Lateral pronotal carina distinct, present only along lower half of posterior margin of anterior pronotal plate. Lateral surface of pronotum foveate-reticulate and densely punctate with conspicuous appressed pubescence. Dorsal pronotal area finely transversely striate, present along anterior fourth of posterior margin of pronotum. Mesoscutum flat dorsally and transversely costate with superficial foveae set in rows and evenly distributed appressed pubescence. Scutellar sulcus divided only by a single median longitudinal carina. Axillar area with conspicuous pubescence. Mesoscutellum foveate-reticulate; mesoscutellum broadly trapezoid posteriorly in dorsal view, with posterior margin distinctly emarginate; lateral dorsal process distinct. Mesopleural triangle conspicuously pubescent and well defined ventrally by a smoothly curved carina. Median mesopleural impression percurrent and glabrous. Upper mesopleuron glabrous and with short pubescence anteriorly; lower mesopleuron glabrous and densely pubescent in ventral impression. Metepisternum areolate-reticulate in upper part and conspicuously pubescent ventrally. Lateral propodeal carina percurrent, strongly curved laterally, and distinctly raised dorsally to form in the middle a strong, broadly rounded process with sparse dorsal pubescence; medi- an propodeal area glabrate, with median longitudinal carina anterior to median transverse carina and two submedian carinae posteriorly. Rs+M of forewing nebulous, arising from middle of basal vein. Marginal cell 3.5 times as long as wide and 1.6 times as long as submarginal cell. Bulla on Sc+R 1 absent.

Petiole 0.60 times as long as wide in lateral view. Tergum 8 completely covered by T7; relative size of abdominal T3–7: 1.6:1.0:1.0:2.1:0.9; T3 glabrous; T4 glabrous with a narrow band of fine punctures across anterior second fifth; T5 glabrous and finely punctate; T6 finely and densely punctate dorsolaterally, with an extensive patch of dense pubescence covering posterior twothirds of upper half of the tergite; and T7 densely punctate, with a broad band of conspicuous pubescence dorsolaterally in the middle and posteriorly glabrous. Apical teeth of metatibia slender and pointed apically. Metatibia with two to three dorsal dents. Apical protuberance of first metatarsomeres reaching slightly beyond middle of second metatarsomere. 1mt (excluding apical protuberance)/2–5mt 5 0.66.

MALE: Unknown.

Paramblynotus nebulosus differs from all other species of the genus except P. miniatus in having its metatibia dorsally with dents. It differs from the latter in (1) forewing with a broad dark gray band along outer margin; (2) upper metepisternum without a large, glabrous elevated median area; (3) head, pronotum, and metasomal T6–7 covered with conspicuous pubescence; and (4) distinctly larger.

TYPE MATERIAL: HOLOTYPE: ♀, Malaysia: NW Sabah, near Long Pa Sia (west), ca. 1,200 m, 1987-IV-2–14, C.v. Arhterberg coll. ( NNMN). PARATYPES: 8♀♀. 1♀, data as holotype ( NNMN) ; 1♀, Indonesia: Sulawesi, near Sangona, Base Camp , Gn. Watuwila, ca. 200 m, 1989-X-10–15, C. v. Achterberg coll. ( NNMN) ; 1♀, Indonesia: Sulawesi, near Mamasa, Penannang , 1,700 m, 1991-IV-10–22, C. v. Achterberg coll. ( NNMN) ; 3♀♀, Malaysia, Negri S., Pasoh Research Station , coll. By P. and M. Becker in 1978-VII-14, 1978-XII-5, and 1979-II-20, respectively ( AEI) ; 1♀, Malaysia: Sabah, Sipitang, Mendolong, T 3/W5, 1989-III-8, S. Adebratt coll. ( ZML) ; 1♀, Indonesia: Sulawesi, Dumoga-BoneTaraut ‘‘ Edwards’ ’, 680 m, 1985-IV-26, J. Martin coll. ( NHM) .

DISTRIBUTION: Malaysia, Borneo; Indonesia, Sulawesi.

ETYMOLOGY: From Latin, nebulosus , meaning misty, foggy, cloudy, or dark, referring to the broad gray band along the outer margin of forewing.