Ectrepesthoneura succinimontana, BLAGODEROV & GRIMALDI, 2004

BLAGODEROV, VLADIMIR & GRIMALDI, DAVID, 2004, Fossil Sciaroidea (Diptera) in Cretaceous Ambers, Exclusive of Cecidomyiidae, Sciaridae, and Keroplatidae, American Museum Novitates 3433 (1), pp. 1-76 : 39

Ectrepesthoneura succinimontana , new species Figures 48, 49 , Plate 5D

DIAGNOSIS: Fore tibiae slightly shorter than fore femur. Section M3 and bases of M 1 and M 2 very weak. Length of small radial cell twice the width.

DESCRIPTION: Body length = 2.39 mm; wing length = 2.02 mm. Head: Occiput and frons with short curved setae. Ocelli three, distance of lateral one to eye margin more than twice the ocellus diameter. Scape subconical, pedicel rounded, width twice flagellomere width. Eleven segments of flagellum preserved, flagellomeres cylindrical, with length about equal to width. Face setose. Palpi 4­segmented; 1st palpomere very small; antepenultimate one oval, its length twice the width, with medial sensory pit; penultimate one with length 3Χ width, attached to the second one preapically; apical segment long and slender. Clypeus triangular, setose. Thorax: Scutum with numerous long setae most­ ly curved forward and shorter ones curved back, arranged in rows. Antepronotum with 3 long setae. Scutellum with two pairs of long setae. Other thoracic sclerites bare. Wing: All longitudinal veins with short setae. Sc meets R proximal to base of M3. Length ratio of sections RS1, RS2, and RS3 is 1:2:13. RS1 transverse, 0.3Χ length of r­m. Crossvein r­m 0.7Χ length of R 1 and 0.5Χ length of M3 section. Costa ends beyond tip of R 5, one­third distance between tips of R 5 and M 1. Section M3 and M 1 and M 2 bases very weak. Legs: Fore coxae with numerous anteromedial setae, hind coxae with posterolateral setae. Fore and mid tibiae slightly shorter than fore femora. Mid and hind tibiae and tarsi with rows of black bristles. Tibial spurs 1.5–2Χ tibial diameter. Tarsal claw with one long tooth. Abdomen setose. Eighth sternite almost triangular. Cerci 2­segmented, basal segment stick­shaped, with length twice the width, extends beyond 8th sternite; apical one discoidal, attached at the apex of the basal one, 2.2Χ shorter. Dorsal border of 8th sternite (gonocoxite 8 of Martinsen and Söli, 2000) straight.

MATERIAL: Holotype PIN 3311 /662, male. Russia: Taimyr Peninsula , Yantardakh, coll. 1971.

ETYMOLOGY: From Latin succinum meaning ‘‘amber’’ and montanus meaning ‘‘mountainous’’. Yantardakh means ‘‘amber mountain’’ in Dolgan language.

COMMENTS: Comparision with recent species is difficult, because nongenital characters used for distinguishing species have been considered to be unreliable for this genus ( Martinsen and Söli, 2000). For species where the female genitalia are known the new species is most similar to E. hirta Winnertz.

Figures

Figs. 48–51. Ectrepesthoneura End. 48. E. succinimontana, n.sp., holotype PIN 3311/662. 49. Female genitalia. 50. E. swolenskyi, n.sp., holotype AMNH NJ­824. 51. Male genitalia.