Saigusaia pikei, BLAGODEROV & GRIMALDI, 2004

BLAGODEROV, VLADIMIR & GRIMALDI, DAVID, 2004, Fossil Sciaroidea (Diptera) in Cretaceous Ambers, Exclusive of Cecidomyiidae, Sciaridae, and Keroplatidae, American Museum Novitates 3433 (1), pp. 1-76 : 31-32

Saigusaia pikei , new species

Figure 37 , Plate 4B

DIAGNOSIS: Distinguished from the living species by Sc long, meeting C just before RS base.

DESCRIPTION (see also table 3): Body length = 2.84 mm, wing length = 1.80 mm. Head: Ocelli three, each slightly raised, very close. Eyes pubescent. Vertex and clypeus setulose. Flagellum 14­segmented, flagellomeres cylindrical, with length slightly greater than width. Mouthparts form short proboscis, length is one­half the head height. Only three palpal segments seen: antepenultimate segment swollen, penultimate and apical ones narrower, bacilliform. Length ratio 1:1.7:2.7. Thorax: Antepronotum with two long setae. Scutum with strong, curved lateral, dorsocentral and acrostichal setae. Mediotergite and laterotergites bare. Scutellum with 5 pairs of rather short setae. Metepisternum with two long setae dorsally. Wing: Costa ends beyond tip of R 5, one­third distance between tips of R 5 and M 1. Stem of R, R 1 and R 5 with rows of dorsal and ventral setae. R 1 and R 5 straight. Sc meets C just before RS base. Sc 2 apical, proximal to base of M3. Crossvein r­m 1.5Χ length of RS1 and 0.3Χ length of M3 section. Fork of M 1 and M 2 2.7Χ length of M3. M 1 and M 2 fork campanulate, veins straight, with a few ventral setae apically. Base of M 3+4 and CuA fork proximal to base of M 1 and M 2 fork, M 3+4 and

TABLE 3 Comparison of Boletina , Saigusaia , and Saigusaia pikei n.sp.

CuA curved caudally. Legs: Tibial bristles 2Χ tibial diameter. Tarsal claw with one tooth. Abdomen: Sternite 8 with two triangular, rounded, caudal lobes covered by long setae. Cerci two­segmented. Basal segment large and broad, apical one 0.5Χ the length and 0.5Χ the width of basal one; both setose.

MATERIAL: Holotype TMPD P79.15.7.21, female. Canada: Alberta, Grassy Lake , coll. T. Pike.

ETYMOLOGY: The species epithet is patronymic for Dr. T. Pike, who collected the specimen.

COMMENTS: This species has almost all the features of Saigusaia (table 3, see also Vockeroth, 1980), so despite the long Sc vein its placement in the genus is virtually certain.

Figures

Figs. 35–37. Dziedzickia and Saigusaia. 35. D. nashi, n.sp., holotype AMNH NJ­117a. 36. Male genitalia of the holotype. 37. Saigusaia pikei, n.sp., holotype TMPD P79.15.7.21