Pseudomanota, BLAGODEROV & GRIMALDI, 2004

BLAGODEROV, VLADIMIR & GRIMALDI, DAVID, 2004, Fossil Sciaroidea (Diptera) in Cretaceous Ambers, Exclusive of Cecidomyiidae, Sciaridae, and Keroplatidae, American Museum Novitates 3433 (1), pp. 1-76 : 25

Pseudomanota , new genus

DIAGNOSIS: Infraorbital setae absent. Three palpomeres, penultimate one with long thin apical appendage. Antepronotum wide. Proepisternum rodlike. Laterotergites and mediotergite bare. Sc short, free. Wing membrane with macrotrichia. Base of M 1 and M 2 fork reduced. M 1 reaches wing margin before wing apex. M 3+4 free at the base. Gonostyli simple, slightly curved.

TYPE SPECIES: Pseudomanota perplexa n.sp.

ETYMOLOGY: The genus name is a combination of pseudos (Greek Ψeυ˘ƃoơ, or a lie, a fraud) and the genus name Manota . The name is feminine.

COMMENTS: Some structures, especially the wide quadrate face, long and flexible apical palpomere, short Sc and R 1, horizontal r­m, and reduced venation are like Manotinae . Also, unlike typical Sciophilinae , the new genus has laterotergites and especially the mediotergite bare. Nevertheless, the genus is attributed to the subfamily Sciophilinae based on a long, rodlike proepimeron, the rather distal position of humeral vein, and long and slender gonocoxites (which are not known for any manotines). Within Sciophilinae the new genus might be close to the Azana group of genera ( Matile, 1998), demonstrating reduction of median veins. All of the genera in the group have reduced venation, with CuA simple and not forming a fork with M 3+4. Matile (1998) supposed that M 3+ 4 in this group was reduced completely, while the fork of M 1 and M 2 lost its base. Jugding on the position of vein apices, at least Paratrizygia , Neoaphelomera , and Neotrizygia may have lost M 2, and an incomplete vein between M 1 and CuA may be homologous to M 3+4. The new species has venation more primitive than the genera in the Azana group, preserving all the longitudinal veins, although without bases.