Protognoriste amplicauda, BLAGODEROV & GRIMALDI, 2004

BLAGODEROV, VLADIMIR & GRIMALDI, DAVID, 2004, Fossil Sciaroidea (Diptera) in Cretaceous Ambers, Exclusive of Cecidomyiidae, Sciaridae, and Keroplatidae, American Museum Novitates 3433 (1), pp. 1-76 : 17-19

Protognoriste amplicauda , new species

Figure 15 , Plate 2B

DIAGNOSIS: R 1 length 3Χ that of r­m. Gonocoxite massive. Gonostyli straight, flattened dorsoventrally.

DESCRIPTION: Body length = 1.25 mm; wing length = 1.11 mm. Head: Eyes setulose, setae very short. Ocelli three, equal, almost in straight line, distance of lateral ocellus from eye margin equal to distance from mid ocellus to lateral one. Vertex and frons setose. Scape very small. Pedicel spherical, obscure. Flagellum 14­segmented, flagellomeres cylindrical, lengths about equal to widths, setose. Palpi short, 3­segmented, apical segment rounded, penultimate cylindrical, length 2Χ the width. Face wide, quadrate. Thorax: Scutum with lateral, dorsocentral and acrostichal setae. Scutellum with several long setae. Mediotergite with short trichia. Metepisternum with anterodorsal cleft, height about equal to width. Wing membrane without macrotrichia. Costa ends beyond tip of R 5, at 1/6 the length between tips of R 5 and M 1. Sc short, ends free at the level of RS base. Humeral cross­vein transverse. R 1 short, about 0.4Χ wing length. R 5 about 0.8Χ wing length, almost straight. RS base very short, transverse. Crossvein r­m horizontal, meets M 3+4 and CuA stem, its length 3Χ less than R 1 length. M 1 almost straight, weakened at base. M 2 and M stem absent. Base of M and CuA fork between levels of RS base and R 1 tip. CuA and M curved caudally. Legs: Coxae almost equal in length, hind coxae bare. Tibial setulae in distinct rows. Tibiae, especially fore, short. Tarsal claw with small obtuse tooth. Abdomen as long as wing, with 8 segments, se­ tose. Eighth segment short, retracted. Gonocoxite massive. Gonostyli straight, flattened dorsoventrally.

3+4

3+4

MATERIAL: Holotype PIN 3426 /257, male. Russia: Taimyr Peninsula , Nizhnyaya Agapa, coll. 1973.

ETYMOLOGY: The species epithet is derived from Latin words amplus meaning ‘‘large, distinguished’’ and cauda meaning ‘‘tail’’ in reference to large terminalia of the species.

Figures

Figs. 15–18. Protognoriste, n.gen. 15. P. amplicauda, n.sp., holotype PIN 3426/257. 16. P. goeleti, n.sp., holotype AMNH Bu­406. 17. P. nascifoa, n.sp., holotype AMNH Bu­434. 18. P. nascifoa, male genitalia.