Lebanognoriste prima, BLAGODEROV & GRIMALDI, 2004

BLAGODEROV, VLADIMIR & GRIMALDI, DAVID, 2004, Fossil Sciaroidea (Diptera) in Cretaceous Ambers, Exclusive of Cecidomyiidae, Sciaridae, and Keroplatidae, American Museum Novitates 3433 (1), pp. 1-76 : 14

Lebanognoriste prima , new species

Figures 9–11 , Plate 1E

DIAGNOSIS: As for genus.

DESCRIPTION: Body length = 1.53 mm; wing length = 1.64 mm. Head: Postocciput flat, with several strong setae behind eye margin. Eyes setose. Ocelli not visible. Scape and pedicel twice as wide as flagellomeres, scape with one and pedicel with two apical rows of setae. Flagellum with 14 cylindrical segments, covered by hairs as long as 1/2 flagellomere diameter. Face setulose. Only three apical segments of palpi visible; antepenultimate and penultimate knoblike; apical long, slender, straight, its length approximately the combined length of preceding two palpomeres. Thorax: Mesonotum with few long lateral and scattered shorter setae. Anterior parapsidal suture distinct. Scutellum with 6 long setae and sparse short ones. Laterotergites and mediotergite bare. Wing membrane without macrotrichia, microtrichia not arranged in rows. All longitudinal veins with setae. Costa ends far beyond tip of R 5. Humeral cross­vein transverse. R 1 short, about 0.4Χ wing length. Sc short, ends in R at middle of r­m. R 5 slightly curved caudally. M 1 slightly curved forward at tip. M fork base and M stem absent. Crossvein r­m short, oblique, fused with base of M 3+4. M and CuA stem short, but distinct. M 3+4 almost straight, CuA curved gently back. Legs: Fore tibiae with simple anteroapical depressed area bearing one row of short bristles. Mid tibia with inner spur 1.5Χ as long as outer one, inner hind tibia spur 1.3Χ as long as outer one. Tibial setulae in distinct rows. Tibial spurs 2–3Χ the tibial diameter. Tarsal claw with one tooth. Abdomen short, setose, with 7 visible segments. Genitalia rounded, rotated at 180°. Gonocoxites massive, fused ventrally. Gonostyli hemicylindrical, slightly curved, shovel­like at apex. Ninth tergite length equals the width, with two deep lateral incisions.

MATERIAL: Holotype AMNH JG268 /1, male. Lebanon: near Jezzine, coll. Aftim and Fadi Acra.

ETYMOLOGY: The species epithet is a Latin word primus meaning ‘‘first’’ and is a reference to the oldest find of the family in the fossil record.

Figures

Figs. 7–11. Archaeognoriste, n.gen. and Lebanognoriste, n.gen. 7. A. primitiva, n.sp., holotype AMNH Bu­1539 8. A. primitiva, male genitalia. 9. L. prima, n.sp. holotype AMNH JG268/1. 10. L. prima, male genitalia dorsally. 11. L. prima, male genitalia ventrally.