Heterostegina reticulata mossanensis, Less & Özcan & Papazzoni & Stockar, 2008

Less, György, Özcan, Ercan, Papazzoni, Cesare A. & Stockar, Rudolf, 2008, The middle to late Eocene evolution of nummulitid foraminifer Heterostegina in the Western Tethys, Acta Palaeontologica Polonica 53 (2), pp. 317-350 : 336-338

Heterostegina reticulata mossanensis ssp. nov.

Figs. 14G–R , 15A–C .

1977 Heterostegina helvetica Kaufmann, 1867 ; Hottinger 1977: textfig. 46B, E.

part 1978 Heterostegina reticulata italica ssp. nov.; Herb 1978: 759, 761, text−figs. 36, 38.

part 1993 Heterostegina reticulata Rütimeyer, 1850 ; Papazzoni and Sirotti 1993: 532, 535–536, pl. 1: 2; pl. 2: 3, 4, 5(?), 6(?).

2004 Heterostegina reticulata mossanensis ssp. nov.; Less and Gyalog 2004: pl. (black and white) 2: 3.

2007 Heterostegina reticulata mossanensis ssp. nov.; Özcan et al. 2007: pl. 1: 19.

Derivation of the name: Named after the type locality of the taxon.

Holotype: MÁFI E. 9555 ( Fig. 14G ), a megalospheric specimen split along the equatorial plane.

MÁFI E. 9522 (I). J–M, O–Q. Heterostegina reticulata multifida ( Bieda, 1949) , latest Bartonian, SBZ 18 C. J, K. Puig Aguilera (NE Spain), A−form, equatorial sections, MÁFI E. 9523 (J), MÁFI E. 9524 (K). L, M, O. Verona (N Italy), Monte Cavro 4, A−form, equatorial sections, MÁFI E. 9525 (L), MÁFI E. 9526 (M), MÁFI E. 9527 (O). P, Q. Mossano 2 (N Italy), A−form, equatorial sections, MÁFI E. 9528 (P), MÁFI E. 9529 (Q). N, R, S. Heterostegina reticulata helvetica Kaufmann, 1867 , latest Bartonian, SBZ 18 C. N. Prella 4 ( Switzerland), MCSNL P4/O2−6, A−form, equatorial section. R, S. Úrhida 4 ( Hungary), A−form, equatorial sections, MÁFI E. 9530 (R), MÁFI E. 9531 (S).

Type locality: Mossano (N Italy, Colli Berici), Marne di Priabona, sample Mossano 6.

Type level: Basal Priabonian, SBZ 19 A.

Material.—35 well−preserved megalospheric specimens split along the equatorial plane.

Diagnosis.— Heterostegina reticulata populations with X mean ranging from 1.7 to 2.8.

The equatorial section of A−forms.—The shape of the regularly arranged chamberlets is slightly hexagonal to almost rectangular. For numerical features see Table 5.

Remarks.—This taxon has been preliminary used by Less and Gyalog (2004) for the population from sample Úrhida 10 and by Özcan et al. (2007) for the population from sample Şarköy 4, however with no description, so the taxon is defined formally here. The figures of Hottinger (1977), Herb (1978) and Papazzoni and Sirotti (1993) cited in the above synonymy list illustrate specimens from the very close vicinity of samples Mossano 5 to 7. These populations are re−evaluated in this paper and based on their parameters, they are classified in synonymy with the present subspecies (although only tentatively in the case of specimens from sample Mossano 16 of Papazzoni and Sirotti 1993 whose position could not be unambiguously correlated with our samples— see also at the description of the Mossano section).

Geographic and stratigraphic distribution.—Earliest Priabonian (SBZ 19 A) of Italy (Mossano, samples 5 to 7; Verona, Castel San Felice: Villa Le Are, Villa Devoto and the outcrop of the hairpin bend), Hungary (Úrhida, sample 10, and Turkey (Şarköy, samples 2, 4, and A).

Figures

Fig. 14. Latest Bartonian and early Priabonian Heterostegina from different Western Tethyan localities. A–F. Heterostegina reticulata reticulata Rütimeyer, 1850, latest Bartonian, SBZ 18 C. A–E. Mossano 3 (N Italy), equatorial sections, MÁFI E. 9549, A−form (A), MÁFI E. 9550, A−form (B), MÁFI E. 9551, A−form (C), MÁFI E. 9552, B−form (D), MÁFI E. 9553, A−form (E). F. Vedi (Armenia), MÁFI E. 9554, A−form, equatorial section. G–R. Heterostegina reticulata mossanensis ssp. nov., earliest Priabonian, SBZ 19 A. G–J. Mossano 6, equatorial sections, holotype, MÁFI E. 9555, A−form (G), paratype, MÁFI E. 9556, A−form (H), paratype, MÁFI E. 9557, A−form (I), paratype, MÁFI E. 9558 (J), B−form. K–M. Úrhida 10 (Hungary), Ą

Fig. 15. Priabonian Heterostegina from different Western Tethyan localities. A–C. Heterostegina reticulata mossanensis ssp. nov., earliest Priabonian, SBZ 19 A. A. Verona (N Italy), Castel San Felice, hairpin bend: MÁFI E. 9565, A−form, equatorial section. B, C: Mossano 7 (N Italy), A−form, equatorial sections, MÁFI E. 9566 (B), MÁFI E. 9567 (C). D–K. Heterostegina reticulata italica Herb, 1978. D, H. Mossano 8 (N Italy), late early Priabonian, SBZ 19 B, A−form, equatorial sections, MÁFI E. 9568 (D), MÁFI E. 9569 (H). E–G, I. Noszvaj (Hungary), “middle” Priabonian, SBZ 19/20. E–G. A−form, equatorial sections, MÁFI E. 9570 (E), MÁFI E. 9571 (F), MÁFI E. 9494 (G). I. MÁFI E. 9572, external view. J, K. Possagno 1 (NE Italy), late Priabonian, Ą