Beaconiella pulchra Shcherbakov, 2021

Shcherbakov, D. E., 2021, New Curvicubitidae and Paraknightiidae (Homoptera: Cicadomorpha) from the Triassic of Central Asia, Russian Entomological Journal 30 (2), pp. 129-134 : 130-132

Beaconiella pulchra Shcherbakov , sp.n.

Figs 1–4 .

MATERIAL. Holotype: tegmen PIN 2785 / 3022(3073)±. Paratypes: tegmina PIN 2240 / 4643(4679)±, 2555/2360, 2368(2386)±, 2429, 2908, 2785/3387(3893)±, 3405; hindwing PIN 2555 /2320±;

Dzhailoucho (Madygen, northern area), Leilek District, Batken Region, SW Kyrgyzstan; Madygen Formation, Ladinian–Carnian.

DESCRIPTION. Tegmen 12.9–19.4 mm long, 6.7–9.1 mm wide (19.4 x 7.1 mm in holotype), variation exaggerat- ed by rock distortion; √LW 9.3–11.8 mm (x1.27). Interradial area about as wide as radial area. RP and MA forks distant from nodal line. Proximal MP section (along nodal line) feebly arched, distal MP section (from nodal line to fork) rather long, sigmoidal, straight or arched backwards, MP1 arched forwards. CuA branches sigmoidal, in middle part subparallel to posterior wing margin. RP, M and CuA altogether with 30–35 terminations. Tegmen fuscous, especially at base and anteriorly, with small pale streaks along costal margin, 2–3 larger markings in pterostigmal area, and pale spots at nodus, on M stem, before crossvein im, and at base of CuA branches. Not only the sequence of distal branching and the number of terminations of all main veins are variable (as is usual in polyneurous wings), but also the degree of Sc+ RA fusion (up to completely free Sc in paratypes 2555/2360, 2785/3405) and the structure of the apex of basal cell: M separating from R at or slightly beyond arculus (in smaller tegmina) or fused with CuA at a very short distance in place of arculus (in larger tegmina). Hindwing 17.4 mm long, 10.4 mm wide; RA, RP, M and CuA forked about same level .

COMPARISON. Similar to Beaconiella fennahi and B. multivenata , but in these latter the interradial area is much wider than the radial area, and the RP and MA forks are close to the nodal line. Distinct from incompletely known B. triassica in smaller size and structure of MP and CuA: in the latter species, the proximal MP section is deeper arched, the distal one is very short, and CuA branches converge with the posterior wing margin.

ETYMOLOGY. Latin pulcher (beautiful, lovely).

Figures

Figs 1–8. Beaconiella spp., Madygen, Triassic: 1–4 — B. pulchra sp.n.: 1–2 — holotype tegmen; 3–4 — paratype hindwing PIN 2555/ 2320 (images mirrored): 3 — hindwing; 4 — coupling hooks along costal margin, SEM; 5–8 — B. cincta sp.n., holotype tegmen:5 — tegmen; 6–8 — SEM: 6 — proximal part of tegmen; 7 — nodus (note granules on veins); 8 — CuA branches (note marginal membrane). Scale bars: 1, 3, 5 — 5 mm, 2 — 3 mm, 4 — 0.5 mm, 6 — 2 mm, 7–8 — 1 mm. Рис. 1–8. Beaconiella spp., Мадыген, триас: 1–4 — B. pulchra sp.n.: 1–2 — переднее крыло, голотип; 3–4 — Заднее крыло, паратип ПИН 2555/2320 (перевернуто Зеркально): 3 — Заднее крыло; 4 — сцепочные крючки по костальному краю, СЭМ; 5–8 — B. cincta sp.n., переднее крыло, голотип: 5 — переднее крыло; 6–8 — СЭМ: 6 —проксимальнаЯ часть крыла; 7 — нодус (видны Зёрна на жилках); 8 — ветви CuA (видна периферическаЯ мембрана). Длина масШтабных линеек: 1, 3, 5 — 5 мм, 2 — 3 мм, 4 — 0.5 мм, 6 — 2 мм, 7–8 — 1 мм.