Vladimiretskiana Fedotova et Perkovsky

Fedotova, Zoya A., Vasilenko, Dmitry V. & Perkovsky, Evgeny E., 2024, Vladimiretskia nathani gen. et sp. nov. (Diptera, Cecidomyiidae: Porricondylinae, Asynaptini, Vladimiretskiana subtr. nov.) from Eocene Rovno amber with description of their eggs, Ecologica Montenegrina 72, pp. 146-157 : 148

Subtribe Vladimiretskiana Fedotova et Perkovsky , subtr. nov.

Type genus: Vladimiretskia Fedotova & Perkovsky , gen. nov.

Diagnosis. Female head: broad; eyes very large, located dorso-laterally. Eyes completely fused at occiput. Рalpi 3-segmented with round dark palpiger (figs. 1 b, c; 2 a; 3 d), much shorter than half height and length of head. Antennae more than 2+14 segmented, flagellomeres with distal neck. Basal node of flagellomeres with well-developed circumfilar sensoria. Vein R 5 almost straight, joining C before wing apex, margin of wing interrupted at this point; cell between C, R 1 not sclerotized. R 1 completely fused proximally with R 5; rs absent; r-m strongly curved distally near joining R 1 (figs 1 a, 3 a); CuA and M 4 simple, well developed from base to end. All legs densely covered by setae. Abdomen without swollen tergites and sternites or long setae. Ovipositor lamellae short, with 3-segmented lamellae. Tenth abdominal sternite does not protrude on ventral ovipositor. Abdomen apically curved upward (figs. 1 a, e; 2 d).

Comparison. The new subtribe strongly differs from Asynaptina sensu lato and others of subfamily Porricondylinae s. str. and Winnertziinae s. str. by its straight R 5 joining C before the wing apex and absence rs, and differs from Winnertziinae by its unsclerotized costal cell and flagellomere sensorial filae not multiporous or uniporous. It is similar to the latter by: simple CuA and M 4; cerci with 3-segmented lamellae as in Diallactiini and Winnerziinae s. l.; upward curved ovipositor apex as in other Asynaptina s. str. and in Winnertziinae ; more than 14 flagellomeres as only in subtribe Asynaptina s. str.; and palpi shorter than half height and length of head like some rare Asynaptina s. str. genera and in different tribes of Porricondylinae s. l. and Winnertziinae s. l. Very short palpi among Asynaptini s. l. are known only in the monotypical genera Asycola Spungis, 1991 and Guyasutomyia Plakidas, 2023 ( Spungis, 1991; Plakidas, 2023). The new subtribe additionally differs from Asynaptina by large completely fused eyes; distally well-developed CuA and M 4; smallest palpi length; and cerci with 3- segmented lamellae (not 2-segmented).