Fannia schnusei Stein, 1911

Dominguez, Martha Cecilia, 2007, A taxonomic revision of the Southern South American species of the genus FANNIA Robineau-Desvoidy (Diptera: Fanniidae), Papéis Avulsos de Zoologia (São Paulo) 47 (24), pp. 289-347 : 336-339

Fannia schnusei Stein, 1911

Figs. 1 B-C, 19 A-F

Fannia schnusei Stein, 1911:106 ; 1919:132; Malloch, 1934:205; Séguy, 1937:174; Ortiz, 1946:157; Pont, 1972:6; Albuquerque et al., 1981:10; Carvalho et al., 1993:15; Pont & Carvalho, 1994:236; Pont, 2001:461; Carvalho et al., 2003:18-19.

Diagnosis: Large black bluish species. Frontal vitta at narrowest point 1.5x width of anterior ocellus. Eye densely covered with long setulae. Parafacial bare. Antenna and palpus black. Thorax lacking vittae. Two pra, 1 near spal and 1 near transverse suture. Fore tarsi yellowish white, fore tarsomeres 1-4, strongly expanded and flattened, bearing 1 apical thorn on posterior margin. F2 strongly curved, with a preapical protuberance on anteroventral surface. T 2 curved, constricted at base, with a prominent subbasal protuberance. Mid tarsomere 1 with a strong ventral basal crest, followed by a long seta. C3 bare at apex of posterior surface. F3 normal to thin strongly bent, with a medial ventral protuberance as broad as femoral width, with a preapical tuft of hair-like and curled pv, 2x femoral width. T 3 expanded and flattened at sub apex, anteroventral and ventral surfaces densely covered with very long and curled setae. Wing lightly tinted. Halter black. Abdomen heart-shaped, metallic blue in ground colour covered with dark brown to black pruinescence.

Type material examined: Paralectotype n. Fannia schnusei Chile, Valparaiso, 12-IX-1902 ( SMTD) Other material examined: Argentina: Neuquén 1 o, Pucará, Parque Nacional Lanín , X-1952 / II-1953, col. Schajiovski ( FIML) . V Región 7 o, 7 n, Melipilla, Curacavi , Ligues de Miraflores , 23-XI-2001, col. J. Solervicens ( CEUM) . 2 o, Valparaíso, Quinta Vergara , 23-X-1963, col. N. Hinchin ( CEUM) . 1 o, EL Salto, Valparaíso , 3-I-1963, col. C. Vivar ( CEUM) . 2 o, Valparaíso , El Granizo, 3-I-1964, col. N. Hichins ( CEUM) . 1 o, Santiago, Caleu , 8-XII-1987, col. Cerda ( MNCN) ; 1 n, Santiago, Caleu , 9-IV-1988, col. Cerda ( MNCN) . 1 o, Santiago, Tiltil, Caleu , 26-IX-1992, col. Cerda ( MNCN) . X Región. 1 o, 1 n, Angol , 9-VII-1928 (from decayed squash) ( USNMNH) ; 1 o, Puerto Varas, Llanquihue , 1-XII-1926, cols. Shannon & Del Ponte ( USNMNH) . Arauco 1 o , Isla Mocha , 15/ 19-III-1971, ( CEUC) .

Description

Male: body length 7-7.6 mm.

Head: Frontal vitta covered with brown pruinescence, at narrowest point 1.5x width of anterior ocellus. Fronto-orbital plate dark grey, at uppermost fr, 0.33x width of anterior ocellus. Twenty long fr, 1.4x length of flagellum. Eye densely covered with long setulae, antero-internal facets larger than remaining; pocl of irregular length in first 1/4. Face and parafacial covered with dark grey pruinescence; parafacial at base of flagellum 0.7x width of same and bare. Facial carina reddish-brown. Gena dark grey to black. Scape and pedicel black, flagellum 2.1x as long as wide and covered with dark grey pruinescence. Arista dark brown at base, lighter brown at apex, short pubescent. Palpus dark brown to black, the apex about as wide as base. Lower oral margin light grey, slightly concave and grooveless.

Thorax: Scutum and scutellum metallic blue in ground colour, covered with black pruinescence. Pleuron grey with blue metallic reflections and covered with brown pruinescence; 4 (3)+4 (6) acr, 1 prescutelar acrs; pprn covered with several long setulae; 2 pra, 1 short, almost indistinguishable from ground setulae, near spal and a second, 3x length of first, near transverse suture; prealar region covered with long setulae; 2 prepm surrounded by numerous long setulae.

Legs: Completely black. F1 normal to thin (length/ width = 6.5); 1 row of pd; 5 to 6 rows of hair-like p; 1 row of pv. T 1 with 1 subapical pd; 1 apical v and pv; 1 row of setulose ad and 1 submedial and 1 apical pd. Fore tarsi yellowish white, tarsomeres 1-4, strongly modified, expanded and flattened and bearing 1 apical thorn on posterior margin ( Fig. 1B ). F2 strongly curved and with a preapical protuberance on anteroventral surface, 1 row of weak ad; 1 row of a; 1 row of hair-like av, as long as femoral width at base, shorter and hooked at apex; 2 rows of pv, one row long and stout, the other shorter and thinner and a third row of shorter row of pv in apical third, where 3 rows are hooked; 1 row of p. T 2 curved, constricted at base, with a prominent subbasal protuberance; ventral pubescence very dense and thin, 0.5x tibial width; 1 long and curved submedial and 3 long apical (the central seta longer) a; 1 subapical ad; 1 strong and straight apical av-v; 1 submedial and 1 apical pd, both reduced; 1 apical short and stout p; apical pv absent. Mid tarsomere 1 with a strong ventral basal crest, followed by a long seta. C3 bare at apex of posterior surface. F3 normal to thin (length/width = 9.8), strongly curved, with a medial ventral protuberance as broad as femoral width; 1 row of ad, longer, stouter and dorsally directed towards apex; 1 row of av, interrupted preapically and 1 av at apex; a preapical tuft of hair-like and curled pv, 2x width of femur ( Fig. 1C ). T 3 expanded and flattened at subapex; 1 submedial and 1 subapical d; 1 submedial and 1 apical ad, lacking medial row of ad; anteroventral and ventral surfaces densely covered with very long and curled setae ( Fig. 1C ).

Wing: Clear, lightly tinted. Lower calypter rounded, upper oval, both white with brown outer margins. Knob of halter black, stalk and base dark brown.

Abdomen: Heart-shaped, tergite 2 more than 2x width of tergite 4; metallic blue in ground colour covered with dark brown to black pruinosity; tergites 2-4, darker in central longitudinal line and posterior margins. Hypopygium not protruding.

Postabdomen: Posterior margin of sternite 5 profoundly indented, and bearing most of setae ( Fig. 19A ). Hypandrial arms directed outwards; postgonite inconspicuous; aedeagus membranous ( Fig. 19B ). Epandrium strongly broadened on posterior region, bell-shaped; cercal plate fused, tapering towards anterior region; bacilliform process cork-screw shaped; surstylus medial, long, broad at base and strongly clavate, weakly connected to epandrium ( Figs. 19 C-D).

Female: body length 6-6.5 mm. Differs from male as follows:

Head: Ocellar triangle short, extending up to second fr. Frontal vitta light brown to reddish-brown. Frons and vitta broad, distance between eye margins wider than 0.33x head width. Ten to 12 fr, 7 longer than flagellum. Upper and lower orb present, with 3 to 4 rows of orbital setulae. Eye sparsely pubescent; pocl of irregular length throughout. Facial carina and gena reddish-brown.

Legs: Fore tarsi unmodified in shape and colour, black; fore tarsomere 1 less than 0.33 length of T 1, tarsomere 4 as long as broad. F2 with 3 rows of a; lacking rows of av and pv. T 2 cone-shaped, narrow at base and broadened towards apex, ventral pubescence absent; 1 apical pv; 1 apical p. Mid tarsomere 1 lacking basal spine and seta on ventral margin. F3 lacking medial posteroventral protuberance; 1 short row of av, 1 subapical, and 1 apical av; lacking subapical tuft of pv. T 3 not modified in shape and lacking long and curled setae on anteroventral surface, with 2 to 3 medial av.

Postabdomen: Cerci normal, short; anal plate longer than broad and covered with setulae; sternite 8 reduced to a pair of circular posterior plates bearing 2 setae and 4 to 5 setulae; post abdominal spiracles 7 and 8 on tergite 6 ( Fig. 19E ). Two pear-shaped spermathecae, slightly grooved, with partially sclerotized ducts ( Fig 19F ).

Distribution: ANDEAN: Southern Chile and Argentina.

Figures

FIGURE 1: F. coxata Shannon & Del ponte. A. Male hind coxa and abdomen, lateral. F. schnusei Stein. B. Male fore tarsi, dorsal. C. Male hind leg, anterior. F. fusconotata (Rondani). D. Male head, anterior. F. scalaris (Fabricius). E. Male mid coxa, anterior. F. roigi sp. nov. F. Male mid tibia, posterior. G. Male hind leg, anterior. F. hermani sp nov. H. Wing.

FIGURE 19: Fannia schnusei Stein. A. Male sternite 5, ventral. B. Male hypandrium, ventral. C. Male terminalia external structures, ventral. D. Male terminalia external structures, lateral. E. Female terminalia external structures. F. Spermathecae.