Thraulodes niger

Kluge, Nikita J., 2020, Systematic position of Thraulodes Ulmer 1920 (Ephemeroptera: Leptophlebiidae) and descriptions of new and little-known species, Zootaxa 4756 (1), pp. 1-142 : 98

Group niger

Definition. In imago and subimago of both sexes, tibiae of all legs blackish-brown, either entirely, or at least in proximal half ( Figs 436–437 , 467–469 , 491–492 , 526–527 ). This coloration is hypodermal and appears in larva of last instar prior to developing imaginal hypodermal coloration on femora.

Composition. Th. niger sp. n., Th. nigrabdominalis sp. n., Th. nigripes sp. n. and Th. nigrotibialis sp. n.

Figures

FIGURES 427–438. Thraulodes niger sp. n. 427–428, head and thorax of male imago in lateral and dorsal view; 429, abdominal terga and sterna of male imago; 430, the same of male subimago; 431, subimaginal mesonotum; 432–433, genitalia of male imago with focus on styliger and on penis; 434, female imago; 435, its left fore wing; 436–437, its fore and middle legs; 438, subimaginal exuviae of fore wing (427–429, 432–433, 438, holotype).

FIGURES 464–473. Thraulodes nigrabdominalis sp. n. (holotype). 464, left half of subimaginal exuviae of mesonotum; 465, 466, male imago; 467–469, fore, middle and hind legs of male imago; 470, subimaginal exuviae of fore leg; 471, genitalia of male imago; 472, apex of penis; 473, subimaginal exuviae of tarsus.

FIGURES 491–497. Thraulodes nigripes sp. n. 491–492, fore and middle legs of female imago; 493–494, female imago in dorsal and lateral view; 495, abdominal terga and sterna of male subimago extracted from larva; 496, genitalia of male subimago extracted from larva; 497, right half of subimaginal exuviae of mesonotum (495–496, holotype).

FIGURES 522–531. Thraulodes nigrotibialis sp. n. 522, right half of subimaginal exuviae of mesonotum; 523, head and thorax of male imago in lateral view; 524–525, female imago in dorsal and lateral view; 526–527, fore and middle leg of male imago; 528, abdominal terga and sterna of male imago; 529–530, genitalia of male imago with focus on styliger and on penis; 531, genitalia of male subimago extracted from larva (522–523, 526–530, holotype).