Bolitocharina Thomson, 1859

Asenjo, Angélico, Irmler, Ulrich, Klimaszewski, Jan, Chandler, Donald S., Fierros-López, Hugo Eduardo & Vieira, Juliana S., 2019, Staphylinidae (Insecta: Coleoptera) in Latin America: synopsis, annotated catalog, diversity and distribution, Zootaxa 4621 (1), pp. 1-406 : 134-140

Subtribe Bolitocharina Thomson, 1859

0445. Bolitochara Mannerheim, 1830: 75

Type species: Aleochara pulchra Gravenhorst, 1806

Agaribiota Bierig, 1937b: 279 [synonymized by Blackwelder 1952: 41]

Type species: Bolitochara cinctigastra Bierig, 1937

Ditropalia Casey, 1906: 263 [synonymized by Seevers 1978: 164]

Type species: Bolitochara bella Märkel, 1844

3 species are included in this genus from the West Indies and South America.

Characters/Description: Fenyes 1920: 114 (characters as Ditropalia ); Coiffait & Sáiz 1967: 65 (characters in key), 79 (characters); Ashe 1992: 365 (characters in key), 367 (characters).

Key: Coiffait & Sáiz 1967: 80 (Chilean species).

Distribution.

West Indies: Cuba.

South America: Colombia, Argentina.

Other regions: Palaearctic, Oriental, Australian, Oceanic.

0446. Habramicrada Pace, 2007: 225 ( Fig. 58 )

Type species: Habramicrada boliviensis Pace, 2007

4 species are known from and are restricted to South America.

Distribution.

South America: Peru, Bolivia, Brazil.

0447. Leptusa Kraatz, 1856b: 60

Type species: Bolitochara pulchella Mannerheim, 1830

Subgenera:

Adexiopisalia Pace, 1982c: 585

Type species: Leptusa hummleri Bernhauer, 1910

Adoxopisalia Pace, 1989: 130

Type species: Leptusa opaca Casey, 1893

Afroleptusa Pace, 1983c: 62

Type species: Leptusa sparsa Fauvel, 1898

Akratopisalia Pace, 1996a: 29

Type species: Leptusa cribrata Pace, 1996

Amalopisalia Pace, 1983c: 97

Type species: Leptusa janetscheki Scheerpeltz, 1956

Anisopisalia Pace, 1989: 214

Type species: Leptusa luigionii Bernhauer, 1935

Anomopisalia Pace, 1983c: 82

Type species: Leptusa granulicauda Eppelsheim, 1890

Anosiopisalia Pace, 1995: 247

Type species: Leptusa nemoricultrix Pace, 1995

Aphaireleptusa Pace, 1996a: 30

Type species: Leptusa anmashanensis Pace, 1996

Aleteleptusa Pace, 1997a: 753 [ Smetana 2004: 429]

Type species: Leptusa chinensis Pace, 1997

Mimumenepisalia Pace, 1997a: 752 [ Smetana 2004: 429]

Type species: Leptusa gonggamontis Pace, 1997

Baldopisalia Pace, 1983e: 130

Type species: Leptusa veronensis Pace, 1983

Boreoleptusa Pace, 1989: 82

Type species: Leptusa norvegica Strand, 1941

Bothrydiopisalia Scheerpeltz, 1966b: 20

Type species: Leptusa bosnica Eppelsheim, 1892

Bryopisalia Pace, 1989: 239

Type species: Leptusa pilatensis Scheerpeltz, 1972

Chondrelytropisalia Scheerpeltz, 1976c: 55

Type species: Leptusa nepalica Scheerpeltz, 1976

Chondridiopisalia Pace, 1983c: 85

Type species: Leptusa granulifera Pace, 1983

Chondridiopisalia Scheerpeltz, 1966b: 19 [nomen nudum, Newton & Thayer 2003]

Type species: Leptusa granulifera Scheerpeltz, 1966

Chondridiopisalia Lohse, 1974: 53 [ Newton & Thayer 2003]

Chondropisalia Scheerpeltz, 1966b: 14

Type species: Leptusa schaschli Ganglbauer, 1897

Cidnopisalia Pace, 1989: 235

Type species: Leptusa cavallensis Pace, 1980

Comphopisalia Scheerpeltz, 1966b: 16

Type species: Leptusa brucki Scriba, 1867

Craspedopisalia Scheerpeltz, 1966b: 19

Type species: Leptusa carinata Bernhauer, 1909

Cryptopisalia Pace, 1983c: 88

Type species: Leptusa montisparii Scheerpeltz, 1953

Cryptopisalia Pace, 1983e: 129 [ Newton & Thayer 2003]

Cyllopisalia Pace, 1982a: 40

Type species: Leptusa difformis Mulsant & Rey, 1853

Cyllopisalia Pace, 1983c: 79 [ Smetana 2004: 431]

Type species: Leptusa difformis Mulsant & Rey, 1853

Parapisalia Scheerpeltz, 1966b: 18 [synonymized by Pace 1982a: 40]

Type species: Leptusa difformis Mulsant & Rey, 1853

Sipalia Mulsant & Rey, 1853b: 32 [synonymized by Pace 1982a: 40]

Type species: Homalota difformis Mulsant & Rey, 1853

Dendroleptusa Pace, 1983c: 65

Type species: Leptusa fumida Erichson, 1839

Drepanoleptusa Pace, 1982d: 93

Type species: Leptusa annapurnensis Pace, 1982

Dysleptusa Pace, 1982c: 580

Type species: Oxypoda fuliginosa Aubé, 1850

Ectinopisalia Scheerpeltz, 1966b: 18

Type species: Leptusa baldensis Ganglbauer, 1895

Emmelopisalia Pace, 1983c: 92

Type species: Leptusa norica Pace, 1979

Entomophallopisalia Pace, 1983c: 90

Type species: Leptusa asturiensis Eppelsheim, 1880

Eospisalia Pace, 1982c: 583

Type species: Leptusa ishizuchiensis Pace, 1982

Eremopisalia Pace, 1989: 196

Type species: Leptusa marianii Pace, 1978

Eucryptusa Casey, 1906: 345

Type species: Silusa nanula Casey, 1893

Evaniopisalia Pace, 1982a: 38

Type species: Leptusa ultracollensis Pace, 1982

Exaeretopisalia Pace, 1989: 233

Type species: Leptusa cansigliensis Pace, 1975

Gallopisalia Pace, 1989: 179

Type species: Leptusa lapidicola Brisout de Barneville, 1863

Gatineauleptusa Klimaszewski in Klimaszewski et al., 2018: 363

Type species: Leptusa gatineauensis Klimaszewski & Pelletier in Klimaszewski et al., 2004

Geopisalia Pace, 1984: 214

Type species: Leptusa peyerimhoffi Bernhauer, 1935

Gnopheropisalia Pace, 1983c: 76

Type species: Leptusa granulipennis Eppelsheim, 1880

Habropisalia Pace, 1989: 216

Type species: Leptusa laticeps Scheerpeltz, 1972

Halmaeusa Kiesenwetter in Kiesenwetter & Kirsch, 1877: 160 [*]

Type species: Halmaeusa antarctica Kiesenwetter, 1877

Antarctophytosus Enderlein, 1909: 377 [synonymized by Steel 1964: 366]

Type species: Phytosus atriceps Waterhouse, 1875

Paraphytosus Cameron, 1917c: 125 [synonymized by Steel 1964: 366]

Type species: Phytosus atriceps Waterhouse, 1875

Austromalota Brèthes, 1925: 170 [synonymized by Steel 1964: 366]

Type species: Austromalota rufimixta Brèthes, 1925

Hemipasilia Pace, 1989: 90

Type species: Leptusa oertzeni Eppelsheim, 1888

Heteroleptusa Pace, 1989: 160

Type species: Sipalia frontalis Casey, 1893

Heterotyphlopasilia Pace, 1983c: 66

Type species: Leptusa serbica Scheerpeltz, 1935

Homopisalia Pace, 1982c: 584

Type species: Leptusa bengalensis Pace, 1982

Ischnopisalia Pace, 1989: 226

Type species: Leptusa poggii Pace, 1978

Kochliodepisalia Pace, 1996a: 35

Type species: Leptusa spirarum Pace, 1996

Koilokoryphepisalia Pace, 1999a: 210

Type species: Leptusa argentierensis Pace, 1999

Lasiopisalia Pace, 1983c: 76

Type species: Leptusa crenulata Bernhauer, 1899

Lathripisalia Pace, 1983e: 135

Type species: Leptusa montispasubii Pace, 1975

Leptopasilia Scheerpeltz, 1966b: 13

Type species: Leptusa hopffgarteni Eppelsheim, 1881

Leptusa Kraatz, 1856b: 60

Type species: Bolitochara pulchella Mannerheim, 1830

Sipalia Mulsant & Rey, 1853a: 45 [ Newton & Thayer 2003]

Liopisalia Pace, 1989: 177

Type species: Leptusa serullazi Sainte-Claire Deville, 1919

Megacolypisalia Pace, 1983e: 133

Type species: Leptusa laevicauda Scheerpeltz, 1958

Melopisalia Pace, 1983c: 88

Type species: Leptusa pasubiana Scheerpeltz, 1953

Metaplasmopasilia Pace, 1989: 104

Type species: Leptusa apennina Holdhaus, 1924

Microcolypisalia Pace, 1989: 196

Type species: Leptusa occulta Pace, 1975

Micropisalia Scheerpeltz, 1948: 164

Type species: Leptusa alpestris Scheerpeltz, 1935

Myopopisalia Scheerpeltz, 1966b: 14

Type species: Leptusa bonvouloiri Brisout de Barneville, 1867

Nanoglossa Fauvel, 1868a: 379 [*]

Type species: Microglossa chilensis Fauvel, 1866

Microglossa Fauvel, 1866b: 282 [synonymized by Fauvel 1868a: 379]

Type species: Microglossa chilensis Fauvel, 1866

Microglossa Fauvel, 1866a: 33 [ Newton & Thayer 2003]

Type species: Microglossa chilensis Fauvel, 1866

Pareuryusa Scheerpeltz, 1972d: 131 [synonymized by Pace 1989: 110]

Type species: Euryusa parallela Fairmaire & Germain, 1862

Nanopisalia Pace, 1983c: 94

Type species: Homalota piceata Mulsant & Rey, 1853

Micropisalia Scheerpeltz, 1966b: 20 [synonymized by Pace 1983c: 94]

Type species: Homalota piceata Mulsant & Rey, 1853

Nanopisalia Pace, 1983e: 132 [ Newton & Thayer 2003]

Type species: Homalota piceata Mulsant & Rey, 1853

Neopisalia Scheerpeltz, 1966b: 16

Type species: Leptusa microphthalma Reitter, 1887

Nesopisalia Pace, 1992b: 112

Type species: Leptusa centralis Pace, 1992

Stenoleptusa Scheerpeltz, 1966b: 10 [synonymized by Assing 2009c: 1286]

Type species: Bolitochara laeviuscula Hochhuth, 1849

Oligopisalia Scheerpeltz, 1966b: 20

Type species: Leptusa flavicornis Brancsik, 1874

Oncopisalia Pace, 1982c: 589

Type species: Leptusa monachorum Bernhauer, 1912

Oreopisalia Pace, 1983c: 82

Type species: Leptusa hoelzeli Scheerpeltz, 1948

Pachygastropisalia Scheerpeltz, 1966b: 15

Type species: Leptusa lativentris Sharp, 1873

Pachygluta Thomson, 1858: 34

Type species: Oxypoda ruficollis Erichson, 1839

Parapisalia Scheerpeltz, 1948: 159

Type species: Leptusa puellaris Hampe, 1863

Pasilia Mulsant & Rey, 1871: 226

Type species: Homalota nubigena Kiesenwetter, 1861

Pasilia Mulsant & Rey, 1872b: 316 [ Newton & Thayer 2003]

Pasilia Mulsant & Rey, 1873c: 73 [ Newton & Thayer 2003]

Pisalia Mulsant & Rey, 1871: 234

Type species: Homalota globulicollis Mulsant & Rey, 1853

Pisalia Mulsant & Rey, 1872b: 324 [ Newton & Thayer 2003]

Pisalia Mulsant & Rey, 1873c: 73 [ Newton & Thayer 2003]

Tropidiopasilia Scheerpeltz, 1966b: 11 [synonymized by Pace 1989: 190]

Type species: Leptusa tricolor Scriba, 1870

Planepisalia Pace, 1989: 217

Type species: Leptusa kochiana Bernhauer, 1936

Protoleptusa Semenov in Semenov et al., 2001: 156

Type species: Leptusa defuncta Semenov, Perkovsky & Petrenko, 2001

Rhombopisalia Scheerpeltz, 1953: 66

Type species: Leptusa knabli Bernhauer, 1909

Roubaliusa Scheerpeltz, 1966b: 13

Type species: Leptusa storkani Roubal, 1916

Scelopisalia Scheerpeltz, 1966b: 18

Type species: Leptusa gracilipes Krauss, 1899

Scoliophallopisalia Pace, 1989: 232

Type species: Leptusa pascuorum Pace, 1975

Scoliophallopisalia Pace, 1981: 115 [nomen nudum, Newton & Thayer 2003]

Scoliophallopisalia Anonymous, 1984: 249 [ Newton & Thayer 2003]

Type species: Leptusa blesioi Pace, 1981

Scolioplatypisalia Pace, 1989: 86

Type species: Leptusa munelensis Apfelbeck, 1907

Stictopisalia Scheerpeltz, 1966b: 16

Type species: Leptusa eximia Kraatz, 1856

Synpisalia Scheerpeltz, 1966b: 16 [synonymized by Pace 1983c: 56]

Type species: Leptusa caucasica Eppelsheim, 1877

Toxophallopisalia Pace, 1983c: 99

Type species: Leptusa austriaca Scheerpeltz, 1935

Trichopasilia Scheerpeltz, 1966b: 12

Type species: Leptusa major Bernhauer, 1900

Typhlopasilia Ganglbauer, 1895: 278

Type species: Leptusa kaufmanni Ganglbauer, 1895

Ulitusa Casey, 1906: 347

Type species: Ulitusa cribratula Casey, 1906

Yunnaleptusa Assing, 2008b: 256

Type species: Leptusa curvata Assing, 2006

16 species are included in this genus from South America.

Characters/Description: Lynch-Arribálzaga 1884: 87 (characters); Fenyes 1920: 120 (characters), 122 (characters subgenera Typhlosilia and Leptusa ), 123 (characters subgenera Eucryptusa and Pachygluta ), 124 (characters subgenera Oreusa and Pasilia ), 125 (characters subgenera Pisalia ); Pace 1989: 25 (characters), 110 (characters of the subgenus Nanoglossa ), 244 (characters of the subgenus Halmaeusa ).

Key: Bernhauer 1900a: 402 (Subgenera and some species); Fenyes 1920: 121 (subgenera); Pace 1989: 25 (subgenera), 110 (species of the subgenus Nanoglossa ), 246 (species of the subgenus Halmaeusa ); Pace 2001a: 12 (Chilean species).

Distribution.

South America: Chile, Argentina, Falkland Isl.

Other regions: Nearctic, Palaearctic, Ethiopian, Oriental, Australian, Antarctic.

0448. Meoticaphaena Pace, 2015e: 72

Type species: Meotica cordobensis Bernhauer, 1934

Only one species is known from and is restricted to South America.

Distribution.

South America: Argentina.

0449. Phymatura J. Sahlberg, 1876: 85

Type species: Bolitochara brevicollis Kraatz, 1856

Venusa Casey, 1906: 272 [synonymized by Ashe 1992: 362]

Type species: Venusa picta Casey, 1906

3 species are included in this genus from South America.

Characters/Description: Fenyes 1920: 116 (characters); Ashe 1992: 366 (characters in key), 371 (characters). Distribution.

South America: Brazil, Argentina.

Other regions: Nearctic, Palaearctic, Ethiopian, Madagascan, Oriental, Australian.

0450. Prosoponotha Pace, 1999c: 146

Type species: Prosoponotha chilensis Pace, 1999

5 species are known from and are restricted to South America.

Distribution.

South America: Brazil, Chile.

0451. Tanystetha Pace, 2002: 137

Type species: Tanystetha brasiliensis Pace, 2002

Only one species is known from and is restricted to South America.

Distribution.

South America: Brazil. Subtribe Dinardopsina Bernhauer & Scheerpeltz 1926

0452. Dinardopsis Bruch, 1917: 257 ( Fig. 52 )

Type species: Dinardopsis solenopsidicola Bruch, 1917

2 species are known from and are restricted to South America.

Distribution.

South America: Brazil, Argentina.

Figures

FIGURES 52–63. Habitus; Aleocharinae (52–63); 52, after Bruch 1917; 53, Peru: department of Madre de Dios (CEMT); 54, Peru: department of Madre de Dios (CEMT); 55, after Pace 2015e; 56, after Pace 2015a; 57, after Pace 2011b; 58, after Pace 2007; 59, after Pace 2008d; 60, after Pace 2015e; 61, after Pace 2014c; 62, after Pace 2011b; 63, after Pace 2011b; slightly modified. Scale bars: 1 mm.