Xanthopygina Sharp, 1884

Asenjo, Angélico, Irmler, Ulrich, Klimaszewski, Jan, Chandler, Donald S., Fierros-López, Hugo Eduardo & Vieira, Juliana S., 2019, Staphylinidae (Insecta: Coleoptera) in Latin America: synopsis, annotated catalog, diversity and distribution, Zootaxa 4621 (1), pp. 1-406 : 305-312

Subtribe Xanthopygina Sharp, 1884

1010. Darwinilus Chatzimanolis, 2014a: 32 ( Fig. 186 )

Type species: Darwinilus sedarisi Chatzimanolis, 2014

Only one species is known from and is restricted to South America.

Distribution.

South America: Argentina.

1011. Dysanellus Bernhauer, 1911a: 419 ( Fig. 189 )

Type species: Dysanellus bruchi Bernhauer, 1911

2 species are known from and are restricted to South America.

Characters/Description: Scheerpeltz 1965b: 277 (characters); Navarrete-Heredia et al. 2002: 299 (characters in key); Chatzimanolis & Ashe 2009: 3 (characters); Chatzimanolis 2012: 267 (characters); Torres-Rodríguez et al. 2012: 219 (characters), 235 (characters in key); Chatzimanolis 2015c: 17 (characters); Chatzimanolis 2018: 280 (characters).

Distribution.

South America: Brazil, Paraguay, Argentina.

1012. Elecatopselaphus Scheerpeltz, 1972c: 39

Type species: Trigonopselaphus violaceus Sharp, 1876

2 species are known from and are restricted to South America.

Distribution.

South America: Colombia, Peru, Brazil.

1013. Elmas Blackwelder, 1952: 146 ( Fig. 190 )

Type species: Selma modesta Sharp, 1876

Selma Sharp, 1876b: 426 [synonymized by Blackwelder 1952: 146]

Type species: Selma modesta Sharp, 1876

20 species are known from and are restricted to Central and South America.

Characters/Description: Ashe & Chatzimanolis 2003: 3 (characters); Torres-Rodríguez et al. 2012: 220 (characters), 235 (characters in key).

Key: Ashe & Chatzimanolis 2003: 33 (species of the genus); Ashe & Chatzimanolis 2006: 28 (species of the genus).

Distribution.

Central America: Nicaragua, Costa Rica, Panama.

South America: Colombia, Suriname, French Guiana, Ecuador, Peru, Bolivia, Brazil.

1014. Gastrisus Sharp, 1876a: 136 ( Fig. 191 )

Type species: Gastrisus laevigatus Sharp, 1876

15 species are known from and are restricted to Central and South America.

Characters/Description: Navarrete-Heredia et al. 2002: 299 (characters in key); Chatzimanolis & Ashe 2009: 3 (characters).

Distribution.

Central America: Mexico, Costa Rica, Panama.

South America: Colombia, Venezuela, Peru, Brazil, Uruguay, Argentina.

1015. Glenus Kraatz, 1857a: 541

Type species: Staphylinus biplagiatus Perty, 1830

20 species are known from and are restricted to Central and South America.

Characters/Description: Sharp 1876a: 147 (characters); Navarrete-Heredia et al. 2002: 299 (characters in key); Torres-Rodríguez et al. 2012: 221 (characters), 235 (characters in key).

Distribution.

Central America: Mexico, Guatemala, Panama.

South America: Colombia, Brazil, Peru, Bolivia, Paraguay, Argentina.

1016. Haematodes Laporte, 1835: 112

Type species: Haematodes bicolor Laporte, 1835

Platycnemus Nordmann, 1837: 135 [synonymized by Lacordaire 1854: 72]

Type species: Platycnemus lateritius Nordmann, 1837

Platytoma Chevrolat in d’Orbigny, 1847: 263 [synonymized by Lacordaire 1854: 72]

Weiserianum Bernhauer, 1927b: 247 [synonymized by Brunke & Chatzimanolis 2018: 5]

Type species: Weiserianum woltersi Bernhauer, 1927

4 species are known from and are restricted to South America.

Characters/Description: Lynch-Arribálzaga 1884: 114 (characters in key), 115 (characters); Erichson 1839a: 340 (characters); Scheerpeltz 1937c: 340 (characters as Weiserianum ); Chatzimanolis 2016: 285 (characters); Brunke & Chatzimanolis 2018: 5 (characters).

Key: Scheerpeltz 1937c: 351 (species as Weiserianum ); Brunke & Chatzimanolis 2018: 5 (species of the genus).

Distribution.

South America: Venezuela?, Brazil, Paraguay, Uruguay, Argentina.

1017. Isanopus Sharp, 1876a: 141 ( Fig. 196 )

Type species: Isanopus tenuicornis Sharp, 1876

6 species are known from and are restricted to Central and South America.

Characters/Description: Navarrete-Heredia et al. 2002: 299 (characters in key); Chatzimanolis 2008: 1769 (characters).

Key: Chatzimanolis 2008: 1789 (species of the genus).

Distribution.

Central America: Mexico, Guatemala, Honduras, Costa Rica, Panama.

South America: Ecuador, Peru, Bolivia, Brazil.

1018. Nausicotus Sharp, 1884: 368

Type species: Trigonopselaphus opacipennis Sharp, 1876

4 species are known from and are restricted to Central and South America.

Characters/Description: Chatzimanolis & Ashe 2009: 3 (characters).

Distribution.

Central America: Nicaragua, Costa Rica.

South America: Brazil, Uruguay, Argentina.

1019. Nordus Blackwelder, 1952: 262

Type species: Brachydirus xanthocerus Nordmann, 1837

Brachydirus Nordmann, 1837: 131 [synonymized by Erichson 1839a: 353]

Type species: Brachydirus xanthocerus Nordmann, 1837

39 species are known from and are restricted to Central and South America.

Characters/Description: Navarrete-Heredia et al. 2002: 299 (characters in key); Chatzimanolis 2004: 6 (characters); Torres-Rodríguez et al. 2012: 223 (characters), 236 (characters in key); Aballay et al. 2014: 104 (characters in key).

Key: Scheerpeltz 1971a: 94 ( Brachydirus ); Chatzimanolis 2004: 47 (species of the genus); Navarrete-Heredia et al. 2006: 105 (species of the genus).

Distribution.

Central America: Mexico, Guatemala, El Salvador, Nicaragua, Costa Rica, Panama.

South America: Colombia, Venezuela, Guyana, Suriname, French Guiana, Ecuador, Peru, Bolivia, Brazil, Paraguay, Argentina.

1020. Ocyolinus Sharp, 1884: 362

Type species: Ocyolinus rugatus Sharp, 1884

7 species are known from and are restricted to Central and South America.

Characters/Description: Chatzimanolis & Ashe 2009: 3 (characters); Chatzimanolis 2017b: 806 (characters).

Key: Chatzimanolis & Ashe 2009: 22 (species of the genus); Chatzimanolis 2017b: 808 (species of the genus). Distribution.

Central America: Guatemala, Nicaragua, Costa Rica, Panama.

South America: Colombia, Venezuela, Brazil.

1021. Oligotergus Bierig, 1937a: 204

Type species: Philothalpus oculatus Bierig, 1937

20 species are known from and are restricted to Latin America.

Characters/Description: Sharp 1876a: 133 (characters); Navarrete-Heredia et al. 2002: 299 (characters in key); Newton & Peck 2006: 152 (characters in key); Torres-Rodríguez et al. 2012: 224 (characters), 235 (characters in key); Aballay et al. 2014: 104 (characters in key); Chatzimanolis & Caron 2016: 132 (characters).

Key: Márquez et al. 2004: 30 (some Mexican species).

Distribution.

West Indies: Trinidad-Tobago.

Central America: Mexico, Guatemala, Nicaragua, Costa Rica, Panama.

South America: Colombia, Venezuela, Guyana, French Guiana, Galapagos Isl., Peru, Bolivia, Brazil, Argentina.

1022. Paraxenopygus Bernhauer, 1911a: 415

Type species: Paraxenopygus tremolerasi Bernhauer, 1911

3 species are known from and are restricted to South America.

Characters/Description: Torres-Rodríguez et al. 2012: 225 (characters), 235 (characters in key).

Distribution.

South America: Peru, Brazil, Uruguay, Argentina.

1023. Phanolinopsis Scheerpeltz, 1968: 67

Type species: Phanolinus discedens Sharp, 1884

6 species are known from and are restricted to Central America.

Characters/Description: Chatzimanolis 2017: 50 (characters).

Key: Chatzimanolis 2017a: 65 (species of the genus).

Distribution.

Central America: Guatemala, Costa Rica, Panama.

South America: Colombia, Ecuador.

1024. Phanolinus Sharp, 1884: 363

Type species: Phanolinus flohri Sharp, 1884

35 species are known from and are restricted to Central and South America.

Characters/Description: Navarrete-Heredia et al. 2002: 299 (characters in key); Torres-Rodríguez et al. 2012: 226 (characters), 236 (characters in key).

Key: Scheerpeltz 1968: 55 (species of the genus); Navarrete-Heredia 2005: 891 (Mexican species).

Distribution.

Central America: Mexico, Guatemala, Nicaragua, Costa Rica, Panama.

South America: Colombia, Venezuela, Ecuador, Peru, Bolivia, Brazil.

1025. Plociopterus Kraatz, 1857a: 539

Type species: Staphylinus fetialis Erichson, 1839

28 species are known from and are restricted to Central and South America.

Characters/Description: Navarrete-Heredia et al. 2002: 299 (characters in key); Torres-Rodríguez et al. 2012: 229 (characters), 235 (characters in key).

Key: Scheerpeltz 1960: 121 (species of the genus).

Distribution.

Central America: Mexico, Guatemala, Honduras, Nicaragua, Costa Rica, Panama.

South America: Colombia, Venezuela, Guyana, French Guiana, Ecuador, Peru, Bolivia, Brazil, Paraguay.

1026. Scaponopselaphus Scheerpeltz, 1972c: 38

Type species: Trigonopselaphus mutator Sharp, 1876

2 species are known from and are restricted to South America.

Characters/Description: Chatzimanolis 2015c: 3 (characters).

Distribution.

South America: Colombia, Guyana, Suriname, French Guiana, Peru, Brazil.

1027. Scariphaeus Erichson, 1839a: 342

Type species: Velleius luridipennis Runde, 1835

4 species are known from and are restricted to South America.

Characters/Description: Lynch-Arribálzaga 1884: 114 (characters in key), 117 (characters); Scheerpeltz 1936: 531 (characters).

Key: Scheerpeltz 1936: 539 (species of the genus).

Distribution.

South America: Ecuador, Brazil, Paraguay, Argentina.

1028. Smilax Laporte, 1835: 116

Type species: Smilax americanus Laporte, 1835

Cordylaspis Nordmann, 1837: 17 [synonymized by Blackwelder 1952: 107 but previously noted by Lacordaire, 1854: 73]

Type species: Cordylaspis tuberculatus Nordmann, 1837

4 species are known from and are restricted to Latin America. Characters/Description: Erichson 1839a: 339–341 (characters); Navarrete-Heredia et al. 2002: 299 (characters in key); Torres-Rodríguez et al. 2012: 230 (characters), 235 (characters in key); Chatzimanolis 2016: 285

(characters). Key: Scheerpeltz 1937a: 335 (species of the Cordylaspis ); Chatzimanolis 2016: 300 (species of the genus).

Distribution.

West Indies: Trinidad-Tobago.

Central America: Mexico?, Nicaragua, Costa Rica, Panama.

South America: Colombia, Venezuela, Guyana, Suriname, French Guiana, Ecuador, Peru, Bolivia, Brazil, Paraguay, Uruguay, Argentina.

1029. Styngetus Sharp, 1884: 361

Type species: Philonthus viduus Erichson, 1840

16 species are known from and are restricted to Central and South America.

Characters/Description: Navarrete-Heredia et al. 2002: 299 (characters in key); Torres-Rodríguez et al. 2012: 230 (characters), 236 (characters in key).

Key: Navarrete-Heredia 1998: 70 (Mexican species).

Distribution.

Central America: Mexico, Guatemala, Costa Rica, Panama.

South America: Colombia, Venezuela, Peru, Bolivia, Brazil, Paraguay, Argentina.

1030. Terataki Chatzimanolis, 2013: 252 ( Fig. 226 )

Type species: Creophilus badiipennis Nordmann, 1837

4 species are known from and are restricted to South America.

Key: Chatzimanolis 2013: 263 (species of the genus).

Distribution.

South America: Brazil, Paraguay.

1031. Torobus Herman, 2001a: 29

Type species: Staphylinus purpurascens Nordmann, 1837

2 species are known from and are restricted to South America.

Characters/Description: Scheerpeltz 1972c: 37–40 (characters); Chatzimanolis & Ashe 2009: 3–4 (characters); Chatzimanolis 2017a: 50 (characters); Chatzimanolis 2018: 285 (characters).

Distribution.

South America: Brazil, Paraguay, Argentina.

1032. Triacrus Nordmann, 1837: 19

Type species: Triacrus dilatatus Nordmann, 1837

Only one species is known from and is restricted to South America.

Characters/Description: Bernhauer 1931: 84 (characters); Chatzimanolis 2015b: 515 (characters).

Distribution.

South America: Brazil, Paraguay, Argentina.

1033. Tricholinus Bernhauer, 1922a: 15

Type species: Tricholinus pehlkei Bernhauer, 1922

Only one species is known from and is restricted to South America.

Distribution.

South America: Colombia.

1034. Trigonopselaphus Gemminger & Harold , 1 868: 597 ( Fig. 228 )

Type species: Trigonophorus myrtillinus Nordmann, 1837

Lypophemus Bernhauer, 1921a: 74 [synonymized by Strand 1921: 180]

Type species: Staphylinus herculeanus Laporte, 1835

Polyphematiana Strand, 1915: 122 [synonymized by Herman 2001a: 28]

Type species: Staphylinus herculeanus Laporte, 1835

Polyphemus Bernhauer in Bernhauer & Schubert, 1914: 397 [synonymized by Scheerpeltz 1933: 1419] Type species: Staphylinus herculeanus Laporte, 1835

Polyphemus Bernhauer, 1914: 51 [synonymized by Scheerpeltz 1933: 1419]

Type species: Polyphemus banghaasi Bernhauer, 1914

Trigonophorus Nordmann, 1837: 8 [synonymized by Gemminger & Harold 1868: 597]

Type species: Trigonophorus myrtillinus Nordmann, 1837

6 species are known from and are restricted to South America. Characters/Description: Lynch-Arribálzaga 1884: 140 (characters); Bernhauer 1914: 51 (characters); Bernhauer

1921a: 74 (characters); Bernhauer 1931: 84 (characters); Smetana & Davies 2000: 18 (characters);

Chatzimanolis 2015a: 168 (characters). Key: Lynch-Arribálzaga 1884: 141 (species of Buenos Aires province, Argentina); Bernhauer 1921e: 19 (species of the genus); Scheerpeltz 1972c: 40 (species of the genus); Chatzimanolis 2015a: 176 (species of the genus). Distribution. South America: Ecuador, Peru, Brazil, Paraguay, Argentina.

1035. Xanthopygus Kraatz, 1856b: 539

Type species: Staphylinus xanthopygus Nordmann, 1837

Heteropygus Bernhauer, 1906b: 195 [synonymized by Blackwelder 1943: 449]

Type species: Xanthopygus oliveirae Lynch-Arribálzaga, 1884

Lampropygus Sharp, 1884: 346 [synonymized by Blackwelder 1943: 449]

Type species: Staphylinus xanthopygus Nordmann, 1837

40 species are included in this genus from Latin America.

Characters/Description: Moore & Legner 1979: 37 (chracters); Lynch-Arribálzaga 1884: 130 (characters); Hayashi

1997: 488 (characters); Blackwelder 1943: 394 (characters in key), 449 (characters); Navarrete-Heredia et al.

2002: 300 (characters in key); Chatzimanolis 2008: 1769 (characters); Torres-Rodríguez et al. 2012: 231

(characters), 236 (characters in key); Chatzimanolis & Caron 2016: 132 (characters).

Key: Lynch-Arribálzaga 1884: 131 (species of Buenos Aires province, Argentina); Scheerpeltz 1969a: 109 (species of the genus); Navarrete-Heredia 2004: 12 (Mexican species).

Distribution.

West Indies: St. Vincent & the Grenadines, Trinidad-Tobago.

Central America: Mexico, Guatemala, Belize, Honduras, El Salvador, Nicaragua, Costa Rica, Panama.

South America: Colombia, Venezuela, Guyana, French Guiana, Ecuador, Peru, Bolivia, Brazil, Paraguay, Uruguay, Argentina.

Other regions: Nearctic.

1036. Xenopygus Bernhauer, 1906b: 196 ( Figs. 229–230 )

Type species: Philonthus analis Erichson, 1840

Leptodiastemus Bernhauer, 1934d: 215 [synonymized by Chatzimanolis & Caron 2016: 132]

Type species: Dysanellus excellens Bernhauer, 1934

9 species are included in this genus from Latin America.

Characters/Description: Navarrete-Heredia et al. 2002: 299 (characters in key); Caron et al. 2016: 63 (characters).

Key: Irmler 1979: 31 (species of the genus); Irmler 1982a: 206 (species of the genus); Márquez et al. 2004: 30 (Mexican species); Torres-Rodríguez et al. 2012: 233 (characters), 236 (characters in key); Aballay et al. 2014: 104 (characters in key); Caron et al. 2016: 65 (species of the genus); Chatzimanolis & Caron 2016: 140 (Key to the punctatus species group in Xenopygus ).

Distribution.

West Indies: Trinidad-Tobago.

Central America: Mexico, Guatemala, Belize, Honduras, El Salvador, Nicaragua, Costa Rica, Panama.

South America: Colombia, Venezuela, Guyana, Suriname, French Guiana, Ecuador, Peru, Bolivia, Brazil, Paraguay, Argentina.

Other regions: Oceanic.

1037. Zackfalinus Chatzimanolis, 2012: 265 ( Fig. 231 )

Type species: Zackfalinus kitrinos Chatzimanolis, 2012

20 species are known from and are restricted to Latin America.

Characters/Description: Chatzimanolis 2015c: 17 (characters).

Key: Chatzimanolis 2012: 302 (species of the genus).

Distribution.

West Indies: Trinidad-Tobago.

Central America: Mexico, Guatemala. Belize, Nicaragua, Costa Rica, Panama.

South America: Colombia, Venezuela, Suriname, French Guiana, Ecuador, Peru, Bolivia, Brazil, Paraguay.

Figures

FIGURES 184–195. Habitus; Staphylininae (184–195); 184, Peru: department of Lima (MUSM); 185, Brazil: state of Paraná (MUSM); 186, after Chatzimanolis 2014a; 187, after Santiago-Jiménez 2008; 188, after Smetana 1982; 189, after Chatzimanolis 2012; 190, Peru: department of Madre de Dios (CEMT); 191, after Navarrete-Heredia & Márquez 1998; 192, after Klimaszewski et al. 2013; 193, after Márquez 2001; 194, after Erichson 1840; 195, after Márquez 2003; slightly modified. Scale bars: 1 mm.

FIGURES 196–207. Habitus; Staphylininae (196–207); 196, after Chatzimanolis 2008; 197, Venezuela: state of Merida (MUSM); 198, Peru: department of Puno (MEKRB); 199, after Chani-Posse & Solodovnikov 2017; 200, after Busanello & Caron 2016; 201, after Brunke & Solodovnikov 2014b; 202, after Asiain & Márquez 2003; 203, after Asiain & Márquez 2012; 204, after Chani-Posse & Solodovnikov 2017; 205, after Costa-Silva et al. 2018; 206, after Chani-Posse 2014b; 207, after Klimaszewski et al. 2013; slightly modified. Scale bars: 1 mm.

FIGURES 220–231. Habitus; Staphylininae (220–231); 220, Peru: department of Madre de Dios (MUSM); 221, Peru: department of Madre de Dios (MUSM); 222, Peru: department of Madre de Dios (MUSM); 223, Chile: region Metropolitana de Santiago (MUSM); 224, after Erichson 1840; 225, after Solodovnikov 2006; 226, after Chatzimanolis 2013; 227, after Márquez & Asiain 2016; 228, Peru: department of Loreto (CEMT); 229, Peru: department of Junin (MUSM); 230, Peru: department of Madre de Dios (CEMT); 231, after Chatzimanolis 2012; slightly modified. Scale bars: 1 mm.