Harpalus (Pseudoophonus) indicus Bates, 1891

Kataev, Boris M. & Liang, Hongbin, 2015, Taxonomic review of Chinese species of ground beetles of the subgenus Pseudoophonus (genus Harpalus) (Coleoptera: Carabidae), Zootaxa 3920 (1), pp. 1-39 : 27-29

Harpalus (Pseudoophonus) indicus Bates, 1891

(Figs 2–3, 60–67)

Diagnosis. This species is somewhat similar to H. davidi and H. pseudohauserianus in having pronotum with rounded basal angles, glabrous elytra with only two lateral intervals punctate and apical spur of protibia not dentate at margins (Figs 2–3), but is easily distinguished from both species by pale legs and very fine microsculpture consisting of narrow transverse meshes on elytral disc in baso-medial portion. Harpalus indicus is also characterized by shallow lateral depressions of pronotum, fused with somewhat wide and shallow basal foveae, very indistinct humeral denticle, poorly visible only from behind, and metafemur with two to three setigerous pores along posterior margin. Internal sac of medial lobe of aedeagus is with long field " a " in basal half and usually with peculiar folded formation " b " in apical half; either without any sclerotized elements or armed with a medial spiny patch ( Figs 60–67 ). Body length 8.9–13.6 mm.

Remarks. Harpalus indicus is widely distributed from North-Western India and North Pakistan throughout the Himalayan region to southern China, Vietnam and Laos. This species forms two subspecies ( Kataev 2014). In China, only the subspecies H. indicus orientalis Kataev, 2014 is present. The nominotypical subspecies occurs in India, Pakistan, Nepal and Buthan ( Kataev 2001, 2014) and will perhaps be found in southern Xizang.

Figures

FIGURES 60 – 67. Harpalus indicus, median lobe of aedeagus. 60 – 63, H. indicus indicus (Nepal); 64 – 67, H. indicus orientalis (64, Vietnam; 65 – 67, Taiwan). 60, 66, view from the left side; 61, 67, dorsal view; 62 – 65, view from the right side). a, b, folded formations of internal sac. Scale = 1.0 mm.