Stenus, LATREILLE

Puthz, Volker, 2017, Übersicht über die neotropischen Arten der Gattung Stenus LATREILLE mit seitlich ungerandetem Abdomen und gelappten Tarsen (Coleoptera, Staphylinidae) 351. Beitrag zur Kenntnis der Steninen, Linzer biologische Beiträge 49 (1), pp. 749-883 : 753-759

Key to the neotropic Stenus with unmargined abdomen and bilobed tarsi

(FB = forebody; S= sternum; T= tergum)

Some characters, which are useful for identification (f. e. the shape of the posterior outline of tergum 10), occur in different (tentatively) monophyletic groups. Species, which share such characters are found in this key under the name "mixed groups" and within this groups with indication to which (tentatively) monophyletic group the respective species belongs.

1 T 10 ancoriform (figs 116-119, 124), S 9 acute apicolaterally. Pronotum short (PW: PL 0.88-0.96), Legs uniformly light coloured OR blackish/brown – yellowish. Small species: FB 1.4-2.0mm............................................................................... S. arculus- group T 10 otherwise.....................................................................................................................2

2 T 10 spiniform (figs 121-123). Species of small or intermediate size: FB 1.3-1.9mm. Pronotum short (PW: PL 0.91-1.03). Frons densely punctate. ................ S. punctatus -group

- T 10 otherwise.....................................................................................................................3

3 T 10 spade-shaped (figs 128-132) ...............................................................mixed group (1)

- T 10 otherwise.....................................................................................................................4

4 T 10 serrate (figs 134. 142-148, 150) ..................................................................................5 T 10 smooth at posterior margin .........................................................................................7

5 Clypeus with a clypeal brush (figs 6, 7, 9) .......................................... S. cupreus -group p.p. Clypeus without a clypeal brush..........................................................................................6

6 ♂: Legs (mostly) with special characters. Pronotum less long (PW: PL 0.86-1.00). Elytra less long (EW: EL 1.01-1.14). S 9 with a short tooth apicolaterally. ♂. E mostly asymmetrical. FB 1.5-2.5 ..................................................................... S. aeneas -group p.p. Otherwise ....................................................................................................mixed group (2)

7 Clypeus with a clypeal brush (figs 6, 7, 9) ..........................................................................8 Clypeus without a clypeal brush (figs 1-5, 8)....................................................................10

8 (Mostly) with metallic luster ...............................................................................................9 Black without metallic luster. S 9 sexually dimorph OR acute apicolaterally (e. g. figs 98, 103, 104, 112). T 10 ± emarginate at posterior margin (e. g. figs 111, 162-164). Pronotum mostly distinctly longer than broad (PW: PL 0.71-0.87). Elytra mostly longer than broad (EW: EL 0.84-0.97). ♂: E with a flagellum FB 2.0- 3.2mm ............................... .......................................................................................................... S. aculeatu s-group pp. Otherwise ....................................................................................................mixed-group (3)

9 (Mostly) with blueish (greenish) metallic luster. Pronotum densely punctate, much longer than broad (PW: PL 0.71-0.87). S 9 either acute or serrate apicolaterally (figs 169-184, 190, 191, 193-195). T 10 mostly smooth at rounded posterior margin. Pronotum much longer than broad (PW: PL 0.73-0.88). ♂: Legs simple. FB 1.8-2.4mm ................................................................................................................... S. cupreus -group

(Mostly) with brassy luster. Pronotum with longitudinal impunctate areas. S 9 acute apicolaterally. T 10 rounded at smooth posterior margin (exception S. ventralis SHARP with a serrate posterior margin, fig. 152). Pronotum much longer than broad (PW: PL 0,73-0.90). ♂: Legs in most species with special characters (e. g. figs 105, 106). FB 1.9-2.9mm ............................................................................................ S. podagricus -group

10 Frons densely punctate without shiny areas ......................................................................11

- Frons with separate shiny areas.........................................................................................12

11 S 9 serrate apicolaterally. T 10 simple. Pronotum less long (PW: PL 0.88-1.00). Elytra short (EW: EL 0.93-1.00). FB 2.0- 2.5mm .............................................. S. vestigialis -group Otherwise ....................................................................................................mixed-group (4)

12 Pronotum with ± distinct lateral bumps (e. g. figs 37, 38, 43). Black or blueish/violet. Pronotum less long (PW: PL 0.83-0.96). Elytra less long (EW: EL 0.91-0.98). S 9 roundly projecting apicolaterally (figs 102, 160) OR acute, with a long apical brush (fig. 107). FB 1.8-2.9mm ....................................................................... S. denticollis -group Pronotum simple. Black without OR with a more or less distinct brassy shine.................13

13 Black with a ± distinct brassy shine. Legs simple (exception S. femoridens PUTHZ ). Pronotum longer (PW: PL 0.80-0.94). Elytra longer (EW: EL 0.94-1.05). S 9 acute apicolaterally. FB 1.7-2.3mm ................................................................. S. aenescens -group

- Black without a metallic shine ....................................................................mixed-group (5)

Mixed group (1)

1 Larger species, FB 2.0mm ................................................................................................2

- Smaller species, FB ≤2.0mm.............................................................................................14

2 Clypeus with a clypeal brush. Body with a blue metallic hue. ♂: E (fig. 315). FB 2.3- 2.4mm. Bolivia ( S. cupreus -group)................................................... S. harpagonifer PUTHZ Clypeus without a clypeal brush. Body black or with brassy hue .......................................3

3 Elytra trapeziform, interstices of abdominal punctation reticulate ......................................4

- Elytra subquadrate, interstices of abdominal punctation smooth.........................................5

4 ♂: Apical portion of median lobe triangular (entire) (fig. 419). FB 2.1mm. Brazil.............. ........................................................................................................... S. mantiqueira PUTHZ

- ♂: Apical portion of median lobe split (fig. 492). FB 2.0- 2.1mm ......................................... ........................................................................................................ S. artipennis L. BENICK

5 Antennae longer, when reflexed extending beyond the posterior margin of the pronotum .............................................................................................................................6

- Antennae shorter, when reflexed not extending beyond the posterior margin of the pronotum .............................................................................................................................7

6 Pronotum densely punctate throughout. Interstices on T 7 smooth. ♂: E (fig. 500). FB 2.0mm. Brazil........................................................................................ S. jugalis ERICHSON

- Pronotum with some shining patches. Interstices on T 7 reticulate. ♂: E (fig. 258). FB 2.2mm. Mexico.................................................................................. S. cuauhtemoc PUTHZ

7 T 10 curved-spade-shaped (fig. 141). Smaller, ♂: E (fig. 487). FB 2.0mm. Venezuela, Colombia, Ecuador, Peru, Bolivia, Trinidad, Surinam, Guyana, Brazil, Paraguay............... ............................................................................................................... S. parviceps SHARP

- T 10 regularly spade-shaped (e. g. figs 131, 132)................................................................8

8 Elytra with a strong medial bump. ♂: Unknown. FB 2.2-2.4mm. Mexico........................... .......................................................................... (S. denticollis- group) S. bullipennis PUTHZ

- Elytra without a medial bump .............................................................................................9

9 Elytral sculpture coarse, very dense, confluent. Pronotum with a small prominent callus on each side. ♂: E (fig. 498). FB 2.2.- 2.4mm. Mexico......................................................... ............................................................................ ( S. denticollis -group) S. scruposus PUTHZ - Punctation of elytra distinctly delimited, not confluent.....................................................10

10 Frons ± densely punctuate.................................................................................................11

- Frons with distinct shining patches ...................................................................................13

11 T 7 densely reticulate. ♂: (fig. 115, PUTHZ 2014). FB 2.2-2.3mm. Argentina...................... .................................................................. ( S. aenescens -group) S. argentinus BERNHAUER T 7 not reticulate (at most some traces of reticulation posteriorly) ...................................12

12 Larger, head slightly broader than elytra (HW: EW 1.01). ♂: E (fig. 446). FB 2.1- 2.2mm. Paraguay, Argentina................(S. aenescens- group) S. paraguayanus BERNHAUER

- Smaller, head slightly narrower than elytra (HW: EW <0.97). ♂: E (fig. 169, PUTHZ 2014). FB 2.0- 2.1mm. Bolivia.......................( S. aenescens -group) S. spinuliventris PUTHZ

13 Abdominal punctation denser, interstices on T 7 mostly smaller than diameter of punctures. ♂: E (fig. 246). FB 2.3mm. U.S.A. (AR, CA, NM), Mexico............................... ............................................................................................................. S. ardoricola PUTHZ

- Abdominal punctation less dense, interstices on T 7 distinctly larger than diameter of punctures. ♂: E (fig. 254). FB 2.2mm. U.S.A. (AR), Mexico............ S. ritamontium PUTHZ

14 Frons densely punctate without distinctly delimited shining patches ................................15

- Frons with dictinctly delimited shining patches ................................................................19

15 Larger, FB 1.7-1.9mm. Elytra broader than long. ♂ unknown. Argentina ( S. aenescens - group)................................................................................................ S. densepilosus PUTHZ

- Smaller, FB 1.3-1.7mm .....................................................................................................16

16 Head broader (HW: EW>0.85).........................................................................................17

- Head less broad (HW: EW <0.80).....................................................................................18

17 Head narrower (HW: EW <0.85). ♂: E (fig. 252). FB 1.3-1.4mm. U.S. A............................ .......................................................................................................... S. meridionalis CASEY

- Head broader (HW: EW 0.90). ♂: E (fig. 257). FB 1.4-1.5mm. U.S. A .............................. ............................................................................................................ S. compressus PUTHZ

18 With brassy shine. T 10 (about as in fig. 138). ♂ unknown. Peru.... S. colossipennis PUTHZ

- Black without brassy shine. T 10 (fig. 128). ♂: E (fig. 451). Chile............... S. gayi SOLIER

19 Interstices on T 7 distinctly reticulate................................................................................20

- Interstices on T 7 smooth ..................................................................................................22

20 ♂: Apex of median lobe narrowly rounded (fig. 413). FB 1.6-1.7 mm. Brazil..................... ....................................................................................... ( S. aeneas -group) S. scurra PUTHZ

- ♂: Apex of median lobe acute...........................................................................................21

21 ♂: Parameres triangularly broadened in anterior quarter (figs 409, 411). FB 1.6-1.9mm. Argentina, Brazil, Paraguay, Peru...................( S. aeneas -group) S. leptocerus BERNHAUER

- ♂: Parameres triangularly broadened in anterior third (fig. 403). FB 1.7mm. Venezuela, Colombia, Ecuador, Peru, Bolivia, Brazil, (?) Paraguay..................................... .................................................................................... ( S. aeneas -group) S. scurrilis PUTHZ

22 Legs brown. ♂: E (fig. 256). FB 1.6mm, Mexico................................................................. ...................................................................... ( S. punctatus -group) S. chimalpopoca PUTHZ

- Legs yellowish or reddish-yellow, apices of femora may be infuscate .............................23

23 Elytra much broader than long (EW: EL 1.10). ♂: E (fig. 472). FB 1.6mm. Chile. ........... ...................................................................................... S. pertusus FAIRMAIRE & GERMAIN

- Elytra less broad, mostly about as broad as long (EW: EL ≤1.07) ....................................24

24 Larger. FB  1.9mm ...........................................................................................................25

- Smaller. FB ≤ 1.7mm .........................................................................................................26

25 Punctation of elytra denser, interstices distinctly smaller than diameter of punctures. ♂: E (fig. 420). FB 1.8-1.9mm. Brazil, Paraguay, (?) Argentina................................................. .................................................................... ( S. aenescens -group) S. plaumannianus PUTHZ

- Punctation of elytra less dense, interstices often as large or larger than diameter of punctures. ♂: E (fig. 501). FB 1.8mm. Brazil................................. S. subleptocerus PUTHZ

26 Punctation of elytra denser, interstices smaller than diameter of punctures ......................27

- Punctation of elytra less dense, interstices mostly much larger than diameter of punctures. ♂: E (fig. 441). FB 1.6mm. Argentina....................... S. callipennis BERNHAUER

27 Head broader (HW: EW 0.80).........................................................................................28

- Head less broad (HW: EW <0.75).....................................................................................30

28 Punctation of T7 coarser and denser, punctures about as large as basal cross section of antennal segment 3, interstices slightly larger than punctures...........................................29

- Punctation of T7 very fine and sparse, punctures finer than medial eye facets, interstices more than twice as large as punctures. ♂: E (fig. 458). FB 1.9mm. Brazil, French Guiana.......................................................................................... S. intonsus PUTHZ

29 Punctation coarser, punctures of pronotum larger than cross section of antennal segment 3. ♂: E (fig. 488). FB 1.4-1.5mm. Peru, Bolivia........... S. peruvianus BERNHAUER

- Punctation less coarse, punctures of pronotum at most as large as cross section of antennal segment 3. ♂: E (fig. 467). FB 1.4-1.5mm. Brazil......................... S. nebulo PUTHZ

30 ♂: E (fig. 447). FB 1.6mm. Brazil, Venezuela, Surinam, Bolivia...... S. frontalis ERICHSON

- ♂: E (fig. 453). FB 1.5-1.7mm. Costa Rica, Mexico................................... S. inanis SHARP

Mixed group (2)

1 Brachypterous. T 10 serrate and spade-shaped. ♂: E (fig. 496). FB: 1.4mm. Jamaica. ........ ........................................................................................................ S. aeneocupreus PUTHZ

- Macropterous ......................................................................................................................2

2 Abdominal pubescence long and conspicuously erect.........................................................3

- Abdominal pubescence less long and recumbent ................................................................4

3 S 9 serrate apicolaterally (figs 153, 193). ♂: Unknown. FB 1.9mm. Brazil.......................... ................................................................................................. S. brasiliensis SCHEERPELTZ

- S 9 acute apicolaterally. ♂: E (fig. 480). FB 1.7-1.8mm. Brazil. .......................................... ...................................................................................................... S. plaumanniellus PUTHZ

4 Larger species, FB 2.0mm ................................................................. S. aeneas -group p.p. (see key in PUTHZ 1984 (* = not in this key): S. acestes PUTHZ *, S. achates PUTHZ *, S. aeneas PUTHZ , S. amata PUTHZ , S. anchises PUTHZ , S. antenor PUTHZ , S. antillensis L. BENICK , S. ascanius PUTHZ , S. cajeta PUTHZ , S. collaris SHARP , S. creusa PUTHZ , S. hippodamea PUTHZ , S. julus PUTHZ , S. latinus PUTHZ , S. lavinia nov.sp. *, S. memmius PUTHZ *, S. ogloblini BERNHAUER , S. operosus ERICHSON , S. palinurus PUTHZ , S. palmipes PUTHZ *)

- Smaller species, FB <2.0mm...............................................................................................5

5 Smaller, FB ≤ 1.6mm ...........................................................................................................6

- Less small, FB  1.6mm .......................................................................................................9

6 Pronotum densely punctate without distinct shiny patches .................................................7

- Pronotum less densely punctate, with distinct shiny patches...............................................8

7 Abdominal punctation denser, interstices on T 5 smaller than diameter of punctures. ♂: E (fig. 440). FB 1.4-1.5mm. Mexico, Honduras, Costa Rica, Trinidad.. S. addendus SHARP

- Abdominal punctation less dense, interstices on T 5 larger than diameter of punctures. ♂: E (fig. 485). FB 1.5-1.6mm. Brazil............................................. S. nebulosus L. BENICK

8 Antennae shorter, when reflexed not extending beyond the posterior margin of pronotum, penultimate segments nearly as long as broad. T 10 very finely serrate. ♂: E (fig. 397). FB 1.5-1.6mm. Brazil................................. ( S. aeneas -group) S. faitoutei PUTHZ - Antennae longer, when reflexed extending beyond the posterior margin of pronotum, penultimate segments longer than broad. T 10 less finely, distinctly serrate (fig. 142). ♂: E (fig. 470). FB 1.6mm. Brazil....................................... S. parcepunctatus BERNHAUER

9 T 10 serrate and spade-shaped (fig. 150). ♂: E (fig. 475). FB 1.7-2.0mm. Brazil, Argentina........................................................................................................ S. plaumanni L. BENICK

- T 10 not spade-shaped.......................................................................................................10

10 Elytral punctation denser, interstices in sutural third not distinctly or much larger than half diameter of punctures .................................................................................................11

- Elytral punctuation less dense, interstices in sutural third distinctly or much larger than half diameter of punctures .................................................................................................12

11 ♂: E (fig. 390).FB 1.8mm. Brazil................................. ( S. aeneas -group) S. agenor PUTHZ

- ♂: E (fig. 407). FB 1.7. Brazil....................................... ( S. aeneas -group) S. mutgo PUTHZ

12 Punctation of frons less dense, median portion broadly smooth. ♂: E (fig. 405). FB 1.8mm. Brazil, Colombia, Ecuador................................... (S. aeneas- group) S. dido PUTHZ

- Punctation of frons denser, median portion only narrowly smooth. ♂: E (fig. 404). FB <2.0mm. Peru ( S. aeneas -group)........................................................... S. pygmalion PUTHZ

Mixed group (3)

1 Smaller, FB ≤ 2.1mm ...........................................................................................................2

- Larger, FB> 2.3mm .............................................................................................................3

2 Punctation denser, interstices on T 7 smaller than diameter of punctures (fig. 90). Frons flat, very densely punctate, interstices smaller than diameter of punctures. ♂: E (fig. 474). FB 1.9mm. Panama.................................................................... S. pilosivestis nov.sp.

- Punctation less dense, interstices on T 7 larger than diameter of punctures (fig. 86). Frons less densely punctate, interstices on slightly elevated median portion larger than diameter of punctures. ♂: E (fig. 460). FB 2.1mm. Brazil......................... S. lapsus nov.sp.

3 Abdominal tergites reticulate ..............................................................................................4

- Abdominal tergites 3-6 not reticulate. ♂: E (fig. 503). FB 2.4-2.5mm. Brazil...................... ................................................................................................................. S. teres ERICHSON

4 ♂: E (fig. 486). FB 2.4-2.5mm. Brazil.................................................. S. parcus L. BENICK

- ♂: E (fig. 452). FB 2.6-2.7mm. Brazil...................................................... S. illautus PUTHZ

Mixed group (4)

1 Pubescence very long and erect...........................................................................................2

- Pubescence less long, recumbent ........................................................................................3

2 Head much broader than elytra (HW: EW>1.20). ♂: E (fig. 456). FB 2.3mm. Colombia..................................................................................................... S. cactus PUTHZ

- Head distinctly less broad than elytra (HW: EW <0.90). ♂: E (fig. 484). FB 2.1- 2.2mm. Mexico........................................................................................ S. hispidus PUTHZ

3 Larger species, FB 2.0mm ................................................................................................4

- Smaller species, FB ≤ 1.9mm ...............................................................................................8

4 Extremely densely punctate, interstices reticulate. ♂: E (fig. 469). FB 2.5mm. Brazil......... ............................................................................................................... S. paraensis PUTHZ

- Less densely punctate, interstices smooth ...........................................................................5

5 With blueish hue. Pronotum with three shiny patches. ♂: E (fig. 481). FB 2.2-2.4mm. Brazil................................................................................................ S. subfrustratus PUTHZ

- Black without blueish hue. Pronotum very densely punctate, without shiny patches..........6

6 Frons very broad (HW: DE 1.4-1.5)....................................................................................7 - Frons less broad (HW: DE 1.9). Smaller. ♂: E (fig. 454). FB 2.0mm. Mexico.................. ........................................................................................................... S. inexoptatus nov.sp.

7 S 9 serrate apicolaterally (fig. 182). ♂: E (fig. 448). FB 2.1-2.2mm. Ecuador...................... ..................................................................................................................... S. fronto PUTHZ S 9 acute apicolaterally. ♂: E (fig. 461). FB 2.2mm. Brazil.................... S. mathani PUTHZ

8 Legs dark brown to blackish. ♂: E (fig. 439). FB 1.3-1.4mm. Chile.................................... ....................................................................................... S. anthrax FAIRMAIRE & GERMAIN

- Legs reddish yellow to reddish brown.................................................................................9

9 Head broader (HW: EW>0.90..........................................................................................10

- Head less broad (HW: EW ≤0.88).....................................................................................12

10 Apicolateral tooth of S9 longer (fig. 167). ♂: E (fig. 449). FB 1.3-1.4mm. Peru................. ................................................................................................................. S. guagua nov.sp.

- Apicolateral tooth of S 9 shorter .......................................................................................11

11 ♂: E (fig. 463). FB 1.3mm. Brazil......................................................... S. lanceatus PUTHZ

- ♂: E (fig. 464). FB 1.3-1.4mm. Brazil................................................. S. lanceolifer PUTHZ

12 Smaller, FB 1.2-1.4mm. ♂: E (fig. 462). Venezuela............................. S. lancearius PUTHZ

- Larger, FB 1.6-1.9mm .......................................................................................................13

13 Elytra longer (EW: EL 0.86). FB 1.9mm. Brazil.................................... S. vexabilis nov.sp.

- Elytra less long (EW: EL 0.89) .......................................................................................14

14 Head broader (HW: EW 0.87-0.88) ................................................................................15 Head less broad (HW: EW 0.83). ♂: E (fig. 291). FB 1.8mm. Peru..... S. sagittifer PUTHZ

15 Slightly more robust, abdominal punctation denser and coarser. ♂: E (fig. 466) FB 1.7mm. Brazil........................................................................................... S. miles PUTHZ

- Slightly less robust, abdominal punctation less dense and less coarse. ♂: E (fig. 450). FB 1.6mm. Brazil.......................................................................................... S. heres SHARP

Mixed group (5)

1 Pubescence very long and erect. ♂: E (fig. 505). 1.7-1.8mm. Brazil.......... S. wallisi PUTHZ

- Pubescence shorter and recumbent......................................................................................2

2 Larger, FB  1.9mm .............................................................................................................3

- Smaller, FB <1.9mm ...........................................................................................................8

3 Head much narrower than elytra, HW: EW ≤0.80...............................................................4

- Head broader, HW: EW>0.85 ............................................................................................5

4 Legs yellowish, frons with large shiny patches. ♂: E (fig. 414). FB 2.0- 2.1mm. Brazi...... l ........................................................................................ ( S. aeneas -group) S. scylla PUTHZ

- Legs dark brown to blackish, frons without or with indistinct shining patches. ♂: E (fig. 482). FB 1.8-1.9mm. Brazil, Peru......................................................... S. batesi SHARP

5 About apical half of metafemora brownish to blackish .......................................................6

- Only apices of femora slightly infuscate. ♂: E (fig. 458). FB 1.9mm. Brazil, French Guiana...................................................................................................... S. intonsus PUTHZ

6 Pronotum broader, (PW: PL 0.95). ♂: E (fig. 445). FB 2.0- 2.1mm. Colombia.................. .............................................................................................................. S. commilito PUTHZ

- Pronotum less broad (PW: PL>0.83)..................................................................................7

7 ♂: E (fig. 443). FB 1.9mm. Peru.......................................................... S. cincinnifer PUTHZ

- ♂: E (fig. 477). FB 2.1mm. Bolivia,? Brazil.......................................... S. pubipenis PUTHZ

8 Head narrower (HW: EW <0.85) ........................................................................................9

- Head broader (HW: EW>0.85..........................................................................................18 9 Legs predominantly brown, dark brown to blackish .........................................................10

- Legs predominantly reddish or reddish yellow..................................................................14

10 T 7 distinctly reticulate......................................................................................................11

- T 7 not reticulate ...............................................................................................................12

11 ♂: E (fig. 490). FB 1.5-1.6mm. Uruguay, Brazil, Argentina,? Peru..................................... ......................................................................................................... S. speculifrons FAUVEL

- ♂: E (fig. 489). FB 1.4-1.6mm. Chile............................................................ S. saizi PUTHZ

12 Smaller, FB <1.4-1.6mm. ♂: E (fig. 439). Chile............ S. anthrax FAIRMAIRE & GERMAIN

- Larger, FB> 1.7mm ...........................................................................................................13

13 Frons densely punctate, punctation of T 7 dense. ♂: E (fig. 482). FB 1.8-1.9mm. Brazil, Peru................................................................................................... S. batesi SHARP

- Frons sparsely punctate, punctation of T 7 sparse. ♂: E (fig. 476). FB 1.8-1.9mm. Brazil............................................................................... S. sanctaecatharinae BERNHAUER

14 Punctures on T 3 larger than diameter of antennal segment 3 ...........................................15

- Punctures on T 3 at most as large as diameter of antennal segment 3 ...............................16

15 Elytra slightly broader than long (EW: EW 1.02). ♂: E (fig. 479). FB 1.5-1.6mm. Bolivia..................................................................................................... S. spoliatus PUTHZ

- Elytra longer than broad (EW: EL 0.96). ♂: E (fig. 473). FB 1.5-1.6mm. Peru, Bolivia......................................................................................... S. peruvianus BERNHAUER

16 Pronotum longer (PW: PL 0.87). ♂: E (fig. 457). FB 1.5-1.6mm. Brazil........................... ......................................................................................................... S. insuspicabilis PUTHZ

- Pronotum less long (PW: PL 0.95)..................................................................................17

17 Elytral punctation denser, interstices in sutural half smaller than diameter of punctures. ♂: E (fig. 491). FB 1.7mm. Venezuela, Brazil. Bolivia, Paraguay....................................... ................................................................................................................ sellatus L. BENICK

- Elytral punctation less dense, interstices in sutural third larger than diameter of punctures. ♂: E (fig. 471). FB 1.8mm. Brazil............................. S. pauloensis BERNHAUER

18 Abdominal punctation coarser and denser, punctures on tergite 7 larger than basal cross section of antennal segment 3, interstices mostly smaller than diameter of punctures. ♂: E (fig. 442). FB 1.5-1.6mm ........................................................... S. chilensis L. BENICK

- Abdominal punctation less coarse and less dense, punctures on T 7 at most as large as basal cross section of antennal segment 3, interstices distinctly to much larger than diameter of punctures ........................................................................................................19

19 T 8 distinctly reticulate......................................................................................................20

- T 8 not reticulate ...............................................................................................................22

20 Antennal segment 10 about 2x as long as broad................................................................21

- Antennal segment 10 nearly as broad as long. ♂: E (fig. 465). FB 1.6mm. Brazil............... .................................................................................................................. S. lancifer PUTHZ

21 ♂: E (fig. 478). FB 1.7-1.8mm. Brazil, Surinam................................... S. ripicursor PUTHZ

- ♂: E (fig. 444). FB 1.6mm. Brazil........................................................ S. denudatus PUTHZ

22 Frons with 5 distinct shiny patches ...................................................................................23

- Frons with 3 shiny patches. ♂: E (fig. 468). FB 1.7-1.8mm. Brazil..... S. nigripontis PUTHZ

23 ♂: E (fig. 459). FB 1.7-1.8mm. Guyana.................................................... S. ludicer PUTHZ

- ♂: E (fig. 438). FB 1.6mm. Brazil...................................................... S. acunculifer PUTHZ

- ♂: E (fig. 473). FB 1.5-1.6mm. French Guiana ..................................... S. shepardi nov.sp.