Campoplex Gravenhorst, 1829

Han, Yuan-Yuan, Achterberg, Kees Van & Chen, Xue-Xin, 2021, The genus Campoplex Gravenhorst, 1829 (Hymenoptera, Ichneumonidae, Campopleginae) from China, Zootaxa 5066 (1), pp. 1-121 : 4-8

publication ID

https://doi.org/ 10.11646/zootaxa.5066.1.1

publication LSID

lsid:zoobank.org:pub:6B3D85E9-31FE-4D08-8E15-BA8959DD1988

DOI

https://doi.org/10.5281/zenodo.5815272

persistent identifier

https://treatment.plazi.org/id/039687B1-477F-6C2E-9DD1-18200C977C76

treatment provided by

Plazi

scientific name

Campoplex Gravenhorst
status

 

Key to species of Campoplex Gravenhorst View in CoL View at ENA from China

1. Face rugose to rugose-punctate ( Fig. 66E View FIGURE 66 ); pronotum punctate, rugulose-punctate or rugose-punctate dorsally; metanotum rugose to rugose-punctate; mesopleuron punctate to rugose-punctate, punctures dense ( Fig. 66B View FIGURE 66 ); propodeum strongly rugose, area superomedia rugulose to trans-striate ( Fig. 66C View FIGURE 66 )......................................................... 2

-. Face granulose to granulose-punctate ( Fig. 2E View FIGURE 2 ), sometimes punctate; pronotum usually granulose to granulose-punctate dorsally, sometimes punctate dorsally, rarely rugulose or rugulose-punctate, never rugose; metanotum usually granulose; mesopleuron usually granulose, punctures sparse to dense ( Fig. 2B View FIGURE 2 ); propodeum granulose to rugose, area superomedia usually granulose ( Fig. 2C View FIGURE 2 ).................................................................................... 6

2. Clypeus rugose; malar space rugulose; epicnemial carina with a notch-like constriction near fore coxa; fore wing areolet small, shorter than stalk................................................................. C. homonae ( Sonan, 1930) View in CoL

-. Clypeus without rugae ( Fig. 66E View FIGURE 66 ; but rugose-punctate in C. xizangensis sp. nov.); malar space nearly smooth to granulose; epicnemial carina without a notch-like constriction near fore coxa; fore wing areolet large, longer than stalk............. 3

3. Clypeus punctate, truncated apically ( Fig. 66E View FIGURE 66 ); mandible without lamella ( Fig. 66E View FIGURE 66 ); occipital carina reaching hypostomal carina above mandible base; pronotum rugose-punctate dorsally; mesoscutum not rugose in notaulic region ( Fig. 66G View FIGURE 66 ); propodeal median area not depressed ( Fig. 66C View FIGURE 66 ); sixth and seventh metasomal tergites with emarginations medially................................................................................................ C. plicopunctatus sp. nov.

-. Clypeus granulose-punctate or rugose-punctate, slightly arched to arched ( Fig. 82E View FIGURE 82 ); mandible with lamella; occipital carina reaching hypostomal carina at mandible base; pronotum punctate to rugulose-punctate dorsally; mesoscutum rugose in notaulic region; propodeal median area moderately to deeply depressed; sixth and seventh metasomal tergites without emarginations medially............................................................................................ 4

4. Clypeus rugose-punctate ( Fig. 82E View FIGURE 82 ); malar space nearly smooth; frons with median carina absent ( Fig. 82E View FIGURE 82 ); interocellar distance 1.3× ocello-ocular distance ( Fig. 82F View FIGURE 82 ); pronotum rugulose-punctate dorsally; propodeal area superomedia rugulose ( Fig. 82C View FIGURE 82 )............................................................................. C. xizangensis sp. nov.

-. Clypeus granulose-punctate ( Fig. 76E View FIGURE 76 ); malar space granulose; frons with median carina present; interocellar distance 1.8–2.0× ocello-ocular distance; pronotum punctate dorsally; propodeal area superomedia rugose to trans-rugose................. 5

5. Face rugose ( Fig. 76E View FIGURE 76 ); scutellum rugose-punctate; mesopleuron rugose-punctate, striations extending to prepectal carina ( Fig. 76B View FIGURE 76 ); latero-longitudinal carina present ( Fig. 76C View FIGURE 76 ); first metasomal segment with dorso-lateral carina absent ( Fig. 76H View FIGURE 76 ); second metasomal tergite apically and third tergite apico-laterally reddish brown......................... C. strigatus sp. nov.

-. Face rugose-punctate ( Fig. 70E View FIGURE 70 ); scutellum granulose-punctate anteriorly, rugose posteriorly; mesopleuron punctate, striations not extending to prepectal carina ( Fig. 70B View FIGURE 70 ); latero-longitudinal carina absent ( Fig. 70C View FIGURE 70 ); first metasomal segment with dorsolateral carina present ( Fig. 70H View FIGURE 70 ); metasoma entirely black................................ C. pseudostrigatus sp. nov.

6. Propodeal medio-longitudinal carina widely diverging posteriorly and often weak to absent ( Fig. 54C View FIGURE 54 ); area superomedia not well formed, sometimes carina stronger where propodeum strongly rugose to striate; otherwise weakly rugose to granulose.. ................................................................................................... 7

-. Propodeal medio-longitudinal carina distinct and not widely diverging posteriorly, parallel-sided or somewhat parallel-sided ( Fig. 20C View FIGURE 20 ); junction between area superomedia and area petiolaris discernible and area superomedia well formed; propodeum always distinctly granulose............................................................................. 15

7. All propodeal carinae very weakly developed ( Fig. 22C View FIGURE 22 ); area basalis confluent with area superomedia area ( Fig. 22C View FIGURE 22 ); area petiolaris granulose-rugulose ( Fig. 22C View FIGURE 22 ); propodeal median area flat ( Fig. 22C View FIGURE 22 ); fore wing areolet absent ( Fig. 22A View FIGURE 22 )......................................................................................... C. confluentus sp. nov.

-. At least some propodeal carinae distinctly developed ( Fig. 54C View FIGURE 54 ); area basalis separated from area superomedia area; propodeal area petiolaris trans-striate or granulose (except C. obtusoclypeus sp. nov.); median area usually depressed; fore wing areolet present.............................................................................................. 8

8. Clypeus smooth and shiny below; interocellar distance approx. 0.8× ocello-ocular distance ( Fig. 54F View FIGURE 54 ); area petiolaris granulose-rugulose ( Fig. 54C View FIGURE 54 ); propodeal median and lateral longitudinal carinae absent ( Fig. 54C View FIGURE 54 ).... C. obtusoclypeus sp. nov.

-. Clypeus entirely mat; interocellar distance 1.0–1.7× ocello-ocular distance; area petiolaris trans-striate or granulose; propodeal median longitudinal carina absent to weakly developed under costula, but gradually stronger developed apically, lateral longitudinal carina absent or present.......................................................................... 9

9. Malar space 0.8× basal width of mandible ( Fig. 34E View FIGURE 34 ); latero-longitudinal carina absent ( Fig. 34C View FIGURE 34 ); fore wing areolet without stalk ( Fig. 34A View FIGURE 34 ); hind femur blackish brown ( Fig. 33 View FIGURE 33 )....................................... C. grandicella sp. nov.

-. Malar space 0.33–0.6× basal width of mandible; latero-longitudinal carina usually present; fore wing areolet with stalk; hind femur yellowish brown to reddish brown (except C. angustaulacis sp. nov.)...................................... 10

10. Interocellar distance approx. 1.0× ocello-ocular distance and 1.4× distance between median and lateral ocelli ( Fig. 64F View FIGURE 64 ); propodeal latero-longitudinal carina strongly developed ( Fig. 64C View FIGURE 64 ); fore wing areolet emitting 2m-cu vein from its basal part ( Fig. 64A View FIGURE 64 )................................................................................ C. protenus sp. nov.

-. Interocellar distance 0.8–1.7× ocello-ocular distance and 1.3–2.5× distance between median and lateral ocelli; latero-longitudinal carina absent to weakly developed; fore wing areolet emitting 2m-cu vein from its apical part..................... 11

11. Propodeum nearly entirely granulose ( Fig. 12C View FIGURE 12 ); propodeal median area broadly and shallowly depressed ( Fig. 12C View FIGURE 12 ); area petiolaris granulose ( Fig. 12C View FIGURE 12 ); second metasomal tergite 0.6–0.7× as long as first tergite........... C. apacicarinatus sp. nov.

-. Propodeum sculpture coarser, not entirely granulose; propodeal median area nearly flat to narrowly and deeply depressed; area petiolaris trans-striate; second metasomal tergite 0.8–0.9× as long as first tergite.................................. 12

12. Mesopleuron and metapleuron strongly and densely granulose, as remainder of mesosoma, dull; propodeum granulose-rugose with a median groove................................................................................. 13

-. Mesopleuron and metapleuron weakly granulose, weaker than remainder of mesosoma; propodeum rugose to granulose-rugose, median area nearly flat........................................................................... 14

13. Interocellar distance approx. 2.5× distance between median and lateral ocelli ( Fig. 10F View FIGURE 10 ); mesopleuron granulose-punctate ( Fig. 10B View FIGURE 10 ); propodeal median area deeply depressed ( Fig. 10C View FIGURE 10 ); fore and mid coxae brown ( Fig. 9 View FIGURE 9 ); hind femur blackish brown ( Fig. 9 View FIGURE 9 ).............................................................................. C. angustaulacis sp. nov.

-. Interocellar distance approx. 1.5× distance between median and lateral ocelli ( Fig. 56F View FIGURE 56 ); mesopleuron granulose to rugulosepunctate ( Fig. 56B View FIGURE 56 ); propodeal median area moderately depressed ( Fig. 56C View FIGURE 56 ); fore and mid coxae yellow ( Fig. 55 View FIGURE 55 ); hind femur yellowish brown ( Fig. 55 View FIGURE 55 )............................................... C. oriens Maheshwary & Gupta, 1977 View in CoL

14. Face granulose medially and rugulose laterally; clypeus strongly granulose; propodeum area dentipara finely granulose; posterior transverse carina making an angle of 80° with pleural carina................ C. chiuae Maheshwary & Gupta, 1977 View in CoL

-. Face finely granulose medially and mat laterally; clypeus mat granulose, with scattered punctures; propodeum area dentipara rugose; posterior transverse carina making an angle of 60° with pleural carina................. C. sauteri ( Uchida, 1932) View in CoL

15. Face and mesoscutum finely punctate; temple punctate; second metasomal tergite punctate; scape and pedicel yellowish brown; hind femur black.............................................................. C. graphoritae ( Uchida, 1942) View in CoL

-. Face and mesoscutum with different sculpture, not finely punctate; temple usually granulose; second metasomal tergite granulose; scape and pedicel yellowish brown to blackish brown; hind femur yellowish brown to black..................... 16

16. Length of body 3.0– 4.5 mm; trans-striations under tegula usually weak; fore wing areolet absent or present with 3rs-m vein incomplete; tarsal claws pectinate and with very weakly developed teeth........................................ 17

-. Length of body 3.6–8.0 mm; trans-striations under tegula usually normally developed; fore wing areolet present; tarsal claws variable............................................................................................ 21

17. Propodeal area superomedia long and narrow; latero-longitudinal carina absent; area petiolaris trans-striate; nervellus intercepted little below middle............................................................................. 18

-. Propodeal area superomedia normally developed; latero-longitudinal carina present; area petiolaris trans-striate or not; nervellus intercepted at lower 0.1–0.3......................................................................... 19

18. Areolet absent ( Fig. 72A View FIGURE 72 ); malar space approx. 0.6× basal width of mandible ( Fig. 72E View FIGURE 72 ); propodeal area superomedia incompletely formed ( Fig. 72C View FIGURE 72 ); temple not swollen behind eyes ( Fig. 72F View FIGURE 72 ); hind femur yellowish brown ( Fig. 71 View FIGURE 71 )............................................................................................... C. assosae Kittel, 2016

-. Areolet present with 3rs-m vein incomplete; malar space approx. equal to basal width of mandible; area superomedia completely formed; temple swollen behind eyes; hind femur brownish............. C. pygmaeus Gupta & Maheshwary, 1977 View in CoL

19. Occipital carina reaching hypostomal carina at mandible base; external angles of second discal cell acute (60°) ( Fig. 4A View FIGURE 4 ); hind wing with nervellus intercepted at lower 0.1 ( Fig. 4A View FIGURE 4 ); tarsal claws not pectinate; first metasomal segment with dorso-lateral carina and lateral groove ( Fig. 4H View FIGURE 4 ); second tergite approx. 3.0× longer than its apical width............ C. acarus sp. nov.

-. Occipital carina reaching hypostomal carina above mandible base; external angles of second discal cell acute (70°); hind wing with nervellus intercepted at lower 0.2–0.3; tarsal claws pectinate but teeth weak; first metasomal segment without dorso-lateral carina and lateral groove; second tergite 1.0–1.1× longer than its apical width.................................... 20

20. Propodeal area petiolaris weakly rugose ( Fig. 28C View FIGURE 28 ); median area slightly depressed ( Fig. 28C View FIGURE 28 ); costula absent ( Fig. 28C View FIGURE 28 ); hind femur approx. 4.6× longer than wide; hind femur black ( Fig. 27 View FIGURE 27 ); metasoma entirely black ( Fig. 27 View FIGURE 27 )..... C. exareola sp. nov.

-. Propodeal area petiolaris trans-striate ( Fig. 36C View FIGURE 36 ); median area not depressed ( Fig. 36C View FIGURE 36 ); costula present ( Fig. 36C View FIGURE 36 ); hind femur approx. 4.0× longer than wide; hind femur yellowish brown ( Fig. 35 View FIGURE 35 ); metasoma not entirely black ( Fig. 35 View FIGURE 35 )................................................................................................ C. granulosus sp. nov.

21. Clypeus long, sharp apically ( Fig. 46E View FIGURE 46 ); malar space 0.9× basal width of mandible ( Fig. 46E View FIGURE 46 ); temple swollen behind eyes ( Fig. 46F View FIGURE 46 ); trans-striations below tegula short and weak ( Fig. 46B View FIGURE 46 ); mesopleural fovea absent ( Fig. 46B View FIGURE 46 ); area petiolaris granuloserugulose ( Fig. 46C View FIGURE 46 ); medio-longitudinal carina developed below costulae and absent apically ( Fig. 46C View FIGURE 46 ); fore wing areolet emitting 2m-cu from its middle ( Fig. 46A View FIGURE 46 ); external angles of second discal cell vertical ( Fig. 46A View FIGURE 46 ); hind wing with nervellus intercepted slightly below middle ( Fig. 46A View FIGURE 46 ); sixth and seventh metasomal tergites with emarginations medially; tegula black ( Fig. 46G View FIGURE 46 )........................................................................ C. longiclypeus sp. nov.

-. Clypeus short, usually not sharp apically; malar space less than 0.9× basal width of mandible; temple swollen or not swollen behind eyes; trans-striations below tegula short and weak to long and strong; mesopleural fovea present; area petiolaris not granulose-rugulose; medio-longitudinal carina completely developed or weakly developed below costulae and gradually stronger apically; fore wing areolet usually emitting 2m-cu from its apical part; external angles of second discal cell acute; hind wing with nervellus usually intercepted below; sixth and seventh metasomal tergites usually without emarginations medially; tegula usually not black..................................................................................... 22

22. Clypeus finely granulose ( Fig. 18E View FIGURE 18 ); area petiolaris granulose medially and rugose laterally ( Fig. 18C View FIGURE 18 ); propodeal latero-longitudinal carina absent ( Fig. 18C View FIGURE 18 ); fore wing areolet emitting 2m-cu from its basal part ( Fig. 18A View FIGURE 18 ); metasoma from third tergite on strongly compressed............................................................... C. collucatus sp. nov.

-. Clypeus granulose to punctate; area petiolaris not granulose medially and rugose laterally; propodeal latero-longitudinal carina present (except in C. tanae sp. nov.); fore wing areolet usually emitting 2m-cu from its apical part; metasoma from third tergite on moderately compressed or not compressed.............................................................. 23

23. Antenna with first flagellomere as long as second flagellomere; clypeus punctate; frons rugose-punctate ( Fig. 62E View FIGURE 62 ); metanotum shallowly punctate; external angles of second discal cell acute (50°) ( Fig. 62A View FIGURE 62 ); sixth and seventh metasomal tergites with weak emarginations medially............................................................... C. proportionis sp. nov.

-. Antenna with first flagellomere usually longer than second flagellomere; clypeus granulose, granulose-punctate or punctate; frons granulose to rugose-punctate; metanotum granulose to rugose-punctate; external angles of second discal cell acute (60– 75°); sixth and seventh metasomal tergites without emarginations medially (except in C. bazariae ).................... 24

24. Intercocellar distance approx. 2.1× ocello-ocular distance ( Fig. 32F View FIGURE 32 ); occipital carina gradually narrowed dorsally; hind wing with nervellus intercepted above its middle ( Fig. 32A View FIGURE 32 ); thyridium large; mandible except teeth black ( Fig. 32E View FIGURE 32 )........................................................................................... C. grandialphus sp. nov.

-. Intercocellar distance 0.8–1.8× ocello-ocular distance; occipital carina evenly arched; hind wing with nervellus intercepted at or below the middle; thyridium small; mandible except teeth yellowish brown.................................... 25

25. Malar space granulose-punctate ( Fig. 6D View FIGURE 6 ); frons rugose-punctate; propodeal area petiolaris rugulose-punctate ( Fig. 6C View FIGURE 6 ); antenna brown ( Fig. 5 View FIGURE 5 ).............................................................. C. adustantennalis sp. nov.

-. Malar space, frons and propodeal area petiolaris with different sculpture; antenna black (except in C. liuae sp. nov.)...... 26

26. Body length 3.6–4.8 mm; clypeus with apical margin truncated or almost truncated; temple slightly swollen to swollen; hind femur 3.8–4.6× longer than wide; tarsal claws pectinate with weakly developed teeth; thyridium separated from basal margin of tergite by its length................................................................................. 27

-. Body length 3.8–8.0 mm; clypeus with apical margin truncated to arched; temple not swollen; hind femur 4.0–5.0× longer than wide; tarsal claws pectinate with weak to strong teeth; position of thyridium variable............................... 30

27. First flagellomere approx. as long as second flagellomere; pronotum rugulose dorsally; mesopleuron punctate ( Fig. 44B View FIGURE 44 ); propodeal area superomedia smooth and shiny, weakly rugulose-granulose ( Fig. 44C View FIGURE 44 ); propodeal latero-longitudinal carina weak ( Fig. 44C View FIGURE 44 ); fore wing vein 1cu-a distad of M&RS by 0.2 of its length ( Fig. 44A View FIGURE 44 ); third metasomal tergite as long as wide apically............................................................................... C. lobatus sp. nov.

-. First flagellomere 1.2–1.4× longer than second flagellomere; pronotum not rugulose dorsally; mesopleuron granulose, granulose-punctate or rugose-punctate; propodeal area superomedia mat, granulose; propodeal latero-longitudinal carina strong; fore wing vein 1cu-a opposite M&RS; third metasomal tergite 0.6–0.7× as long as its apical width........................ 28

28. Malar space approx. 0.7× basal width of mandible ( Fig. 16E View FIGURE 16 ); mandible with upper tooth as long as its lower tooth ( Fig. 16E View FIGURE 16 ); temple slightly swollen behind eyes ( Fig. 16F View FIGURE 16 ); pronotum rugulose-punctate dorsally................ C. atricrus sp. nov.

-. Malar space approx. 0.5× basal width of mandible; mandible with upper tooth longer than its lower tooth; temple swollen behind eyes; pronotum granulose dorsally................................................................... 29

29. Mesoscutum and scutellum granulose-punctate ( Fig. 24G View FIGURE 24 ); mesopleuron granulose-punctate ( Fig. 24B View FIGURE 24 ); hind wing with nervellus inclivous ( Fig. 24A View FIGURE 24 ); inner spur of hind tibia approx. 0.5× as long as first tarsomere of hind tarsus; dorso-lateral carina and lateral groove of first metasomal segment present ( Fig. 24H View FIGURE 24 ); metasoma entirely black ( Fig. 23 View FIGURE 23 ); mid femur brownish, hind femur blackish brown ( Fig. 23 View FIGURE 23 )............................................................. C. cyclus sp. nov.

-. Mesoscutum and scutellum granulose ( Fig. 68G View FIGURE 68 ); mesopleuron granulose ( Fig. 68B View FIGURE 68 ); hind wing with nervellus vertical ( Fig. 68A View FIGURE 68 ); inner spur of hind tibia approx. 0.6× as long as first tarsomere of hind tarsus; dorso-lateral carina and lateral groove of first metasomal segment absent ( Fig. 68H View FIGURE 68 ); metasoma not entirely black ( Fig. 67 View FIGURE 67 ); mid and hind femur yellowish brown ( Fig. 67 View FIGURE 67 ).............................................................................. C. pseudocyclus sp. nov.

30. Propodeal area petiolaris granulose ( Fig. 78C View FIGURE 78 ); hind wing nervellus vertical ( Fig. 78A View FIGURE 78 ); hind femur blackish brown ( Fig. 77 View FIGURE 77 ); hind tibia clearly banded ( Fig. 77 View FIGURE 77 ).......................................................... C. taenius sp. nov.

-. Propodeal area petiolaris not granulose; hind wing nervellus usually inclivous; hind femur yellowish brown to blackish; colour of hind tibia variable.................................................................................. 31

31. Propodeal area superomedia granulose-rugulose ( Fig. 30C View FIGURE 30 ); fore wing marginal cell with distal part of surrounding vein approx. 2.5× longer than proximal one ( Fig. 30A View FIGURE 30 ); fore wing vein 1cu-a distad of M&RS by approx. 0.3 of its length ( Fig. 30A View FIGURE 30 ); metasoma entirely granulose except petiole; thyridium separated from basal margin of tergite by approx. 0.6× of its length; tegula blackish brown ( Fig. 30G View FIGURE 30 ).............................................................. C. granalvus sp. nov.

-. Propodeal area superomedia usually granulose; fore wing marginal cell with distal part of surrounding vein 1.5–2.0× longer than proximal one; fore wing vein 1cu-a opposite or slightly distad of M&RS; metasoma usually only postpetiole and second tergite granulose; thyridium separated from basal margin of tergite by at least its length; tegula yellow to yellowish brown..32

32. Face finely coriaceous, densely punctate; interocellar area distinctly punctate; second and subsequent metasomal tergites finely coriaceous; sixth and seventh metasomal tergites with emarginations medially.................. C. bazariae Sheng, 2014

-. Face granulose to granulose punctate; interocellar area granulose; second and subsequent metasomal tergites mat to coriaceous; sixth and seventh tergites without emarginations medially.................................................... 33

33. First flagellomere as long as second flagellomere; propodeal area basalis with distinct carina posteriorly ( Fig. 42C View FIGURE 42 ); propodeal area petiolaris granulose, rugose laterally ( Fig. 42C View FIGURE 42 ); antenna brown ( Fig. 41 View FIGURE 41 ); postpetiole yellowish brown ( Fig. 41 View FIGURE 41 ).............................................................................................. C. liuae sp. nov.

-. First flagellomere longer than second flagellomere; propodeal area basalis without carina posteriorly; propodeal area petiolaris usually trans-striate; antenna black; postpetiole black........................................................ 34

34. Malar space smooth; area petiolaris rugulose-granulose ( Fig. 60C View FIGURE 60 ); hind wing with nervellus vertical ( Fig. 60A View FIGURE 60 ); metasoma from third tergite on yellowish brown laterally and remainder of metasoma black ( Fig. 59 View FIGURE 59 )...... C. perpendicularis sp. nov.

-. Malar space sculptured; area petiolaris not rugulose-granulose; hind wing with nervellus usually inclivous; metasoma usually entirely black....................................................................................... 35

35. Metanotum granulose-rugose; propodeal area superomedia rugulose ( Fig. 38C View FIGURE 38 ); hind leg except coxa and telotarsi yellow...................................................................................... C. galbipedis sp. nov.

-. Metanotum not granulose-rugose; propodeal area superomedia not rugulose; hind leg sometimes with femur black, tibia not entirely yellow and tarsus usually wholly blackish brown..................................................... 36

36. Malar space shiny, approx. 0.3× basal width of mandible; mandible with upper tooth longer than lower tooth; basal 0.7 of mid femur blackish brown ( Fig. 13 View FIGURE 13 ).......................................................... C. artivultus sp. nov.

-. Malar space not shiny, 0.45–0.8× basal width of mandible; mandible with upper tooth usually as long as lower tooth; mid femur usually yellowish brown............................................................................... 37

37. Frons with median carina present; metasoma not entirely black................................................ 38

-. Frons with median carina absent; metasoma entirely black.................................................... 40

38. Face granulose-punctate ( Fig. 50E View FIGURE 50 ); metanotum rugose-punctate; thyridium separated from basal margin of tergite by its diameter; fore and mid coxae blackish; metasoma black with second tergite laterally yellowish brown ( Fig. 49 View FIGURE 49 )............................................................................................... C. medicarinatus sp. nov.

-. Face granulose; metanotum granulose; thyridium separated from basal margin of tergite equal or more than its diameter; fore and mid coxae at least apically yellow; metasoma from third tergite on laterally yellowish brown..................... 39

39. Interocellar distance approx. 1.8× ocello-ocular distance and 2.3× distance between median and lateral ocelli ( Fig. 26F View FIGURE 26 ); occipital carina reaching hypostomal carina at mandible base; pronotum punctate dorsally; mesopleuron granulose-punctate with punctures dense, punctures separated less than their diameter ( Fig. 26B View FIGURE 26 ); metapleuron punctate ( Fig. 26B View FIGURE 26 ); propodeal area petiolaris trans-striate ( Fig. 26C View FIGURE 26 ); propodeal latero-longitudinal carina present ( Fig. 26C View FIGURE 26 ).......... C. densipunctatus sp. nov.

-. Interocellar distance approx. equal to ocello-ocular distance and 1.4× distance between median and lateral ocelli ( Fig. 80E View FIGURE 80 ); occipital carina reaching hypostomal carina above mandible base; pronotum granulose dorsally; mesopleuron granulose with punctures sparse, punctures separated large than their diameter ( Fig. 80B View FIGURE 80 ); metapleuron granulose ( Fig. 80B View FIGURE 80 ); propodeal area petiolaris rugose ( Fig. 80C View FIGURE 80 ); propodeal latero-longitudinal carina absent ( Fig. 80C View FIGURE 80 ).................... C. tanae sp. nov.

40. Malar space 0.45× basal width of mandible ( Fig. 2E View FIGURE 2 ); mandible with upper tooth slightly longer than lower tooth; trans-striations below tegula weak ( Fig. 2B View FIGURE 2 ); propodeal area petiolaris rugose ( Fig. 2C View FIGURE 2 ); second metasomal tergite 0.9× as long as first tergite................................................................................ C. absitus sp. nov.

-. Malar space more than 0.45× basal width of mandible; mandible with upper tooth as long as lower tooth; trans-striations below tegula weak to strong; propodeal area petiolaris usually trans-striate; second metasomal tergite usually less than 0.9× first tergite............................................................................................... 41

41. Propodeal area superomedia rugose; fore wing pterostigma whitish basally; areolet emitting 2m-cu vein from its middle part; fore and mid coxae yellow........................................................ C. egregius ( Kokujev, 1915) View in CoL

-. Propodeal area superomedia granulose; fore wing pterostigma usually not whitish basally; areolet emitting 2m-cu vein from its apical part; fore and mid coxae usually black.............................................................. 42

42. Hind femur black; hind tibia clearly banded, yellowish brown medially......................................... 43

-. Hind femur yellowish brown; hind tibia usually not clearly banded, unicolored or infuscated apically.................. 46

43. Interocellar distance approx. 1.3× ocello-ocular distance ( Fig. 48F View FIGURE 48 ); propodeal petiolar area rugose ( Fig. 48C View FIGURE 48 ); all trochanters blackish brown................................................................. C. maurotrochanter sp. nov.

-. Interocellar distance 0.8–1.1× ocello-ocular distance; propodeal petiolar area trans-striate; not all trochanters blackish brown. .................................................................................................. 44

44. Malar space approx. 0.8× basal width of mandible; metapleuron and propodeum coarsely granulose, sculpture different from mesopleuron; middle femur with a basal black ring....................... C. maximalus Gupta & Maheshwary, 1977 View in CoL

-. Malar space 0.5–0.7× basal width of mandible; metapleuron and propodeum with normally granulose, sculpture more or less resembling to that of mesopleuron; middle femur entirely yellowish brown...................................... 45

45. Clypeus with apical margin truncated ( Fig. 58E View FIGURE 58 ); pronotum granulose with sparse punctures dorsally; hind wing with nervellus inclivous ( Fig. 58A View FIGURE 58 ).................................................................. C. parassosae sp. nov.

-. Clypeus with apical margin slightly arched; pronotum rugulose-mat dorsally; hind wing with nervellus vertical......................................................................... C. septentrionalis Gupta & Maheshwary, 1977 View in CoL

46. Clypeus with apical margin truncate ( Fig. 74E View FIGURE 74 ); propodeal area petiolaris strongly rugose ( Fig. 74C View FIGURE 74 ); first metasomal segment without dorso-lateral carina and lateral groove ( Fig. 74H View FIGURE 74 ); fore wing pterostigma whitish basally ( Fig. 74A View FIGURE 74 ); fore and mid trochanters and trochantellus brown....................................................... C. shanxiensis sp. nov.

-. Clypeus with apical margin slightly arched to arched; propodeal area petiolaris trans-striate; first metasomal segment with dorso-lateral carina and lateral groove; fore wing pterostigma not whitish basally; fore and mid trochanters and trochantellus yellow to yellowish brown............................................................................. 47

47. Pronotum punctate dorsally; mesopleuron punctate; hind femur 4.0–4.1× longer than wide.......................... 48

-. Pronotum granulose dorsally; mesopleuron granulose to granulose-punctate; hind femur 4.5–5.0× longer than wide...... 49

48. Frons granulose-punctate ( Fig. 20E View FIGURE 20 ); propodeum compact ( Fig. 20C View FIGURE 20 ); hind wing with nervellus intercepted at lower 0.1 of its length ( Fig. 20A View FIGURE 20 ); thyridium separated from basal margin of tergite by 1.5× its diameter; third tergite approx. 0.6× as long as its apical width; scape and pedicel brown ( Fig. 20E View FIGURE 20 ); fore and mid coxae brown.................... C. concretus sp. nov.

-. Frons granulose; propodeum not compact; hind wing with nervellus intercepted at lower 0.3 of its length ( Fig. 52A View FIGURE 52 ); thyridium separated from basal margin of tergite by its diameter; third tergite approx. 0.8× as long as its apical width; scape and pedicel yellowish brown anteriorly and brown dorsally ( Fig. 52E View FIGURE 52 ); fore and mid coxae yellow............ C. monochroma sp. nov.

49. Propodeal sculpture similar to that on mesopleuron and metapleuron; propodeal area dentipara granulose ( Fig. 40C View FIGURE 40 ); second metasomal tergite approx. 0.6× as long as first tergite.............................................. C. hei sp. nov.

-. Propodeal sculpture coarser to that on mesopleuron and metapleuron; propodeal area dentipara granulose-rugose; second metasomal tergite 0.7–0.9× as long as first tergite............................................................... 50

50. Punctures of mesopleuron large and dense ( Fig. 84B View FIGURE 84 ); all propodeal carinae strong ( Fig. 84C View FIGURE 84 ); malar space yellowish brown ( Fig. 84E View FIGURE 84 )....................................................................... C. xuthomelonus sp. nov.

-. Punctures of mesopleuron small and sparse ( Fig. 8B View FIGURE 8 ); all propodeal carinae relatively weak ( Fig. 8C View FIGURE 8 ); malar space black ( Fig. 8E View FIGURE 8 )............................................................... C. anatolus Gupta & Maheshwary, 1977 View in CoL

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