Dima pelionensis, Mertlik, Josef, Németh, Tamás & Kundrata, Robin, 2017

Mertlik, Josef, Németh, Tamás & Kundrata, Robin, 2017, Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula, Zootaxa 4220 (1), pp. 1-63 : 29-30

Dima pelionensis sp. nov.

Figs 92–95 , 152–153 , 209 , 238 , 245 .

Type material. Holotype: GREECE: distr. Magnissia, Mt. Pilio, Chania env., 39°23'52.96'', 23°3'24.54''E, 1263 m, singled at night from ground, 11.VI.2015, ♂, leg. P. Brůha, J. Mertlik, T. Németh & B. Zbuzek ( HNHM) . Paratypes: GREECE: same data as holotype, 8 ♂♂, 6 ♀♀ ( HNHM) ; dtto, 20 ♂♂, 9 ♀♀ (PCRK); dtto, 19 ♂♂, 18 ♀♀, J. Mertlik leg. (PCJM); dtto, 6 ♂♂, 5 ♀♀, B. Zbuzek leg. (PCBZ); dtto, 2 ♂♂, 3 ♀♀, P. Brůha leg. (PCPB); Agios Onoufrios (Volos), 10.VI.2013, 1 ♂, 1 ♀, M. Malavasi leg. ( PCGP) ; Mt. Pelio, Chania env., 1133 m (N39°23'37", E23°02'39"), 3.VI.2011, 1 ♂, J. Krátký leg. ( PCJM) ; distr. Magnissia, Mt. Pilio, Chania village env., 1263 m (39°23'52.96"N, 23°03'24.54"E), 15.VI.2012, 1 ♂, 1 ♀, M. Samek leg. ( PCMS) ; dtto, 35 ♂♂, 30 ♀♀, J. Mertlik leg. (PCJM); dtto, 1 ♀ (PCRK); dtto, 10 ♂♂, 24 ♀♀, V. Dušánek leg. (PCVD); dtto, 17 ♂♂, 17 ♀♀, P. Brůha leg. (PCPB); dtto, 15 ♂♂, 17 ♀♀, B. Zbuzek leg. (PCBZ); Thessalia, Volos, Mt. Pilion , 1100 m, 7.VI.1985, 1 ♂, 1 ♀, G. Etonti leg. (as D. hladilorum in Schimmel & Platia 2008 ) ( PCGP) ; Magnissia, O. Pilio, 900 m, str. Hánia-Makriráhi , 13.VI.1991 / 7.VI.1992, 2 ♀♀, leg. P. M. Giachino & D. Vailati (paratypes of D. fokidensis ) ( PCGP, PCRS) .

Diagnosis. Dima pelionensis is a medium-sized species (body length: 10.5–14.0 mm), with sparsely punctate shiny pronotum with sides bearing decumbent pubescence in the anterior two thirds and semi-erect setae in the posterior third ( Figs 152–153 ). Dima fthiotidensis and D. evritaniensis differ from D. pelionensis by the denser elytral pubescence ( Figs 26–28 , 35–41 ), pronotum matt, with denser and finer punctation ( Figs 126–127, 130– 134 ), and more or less convex scutellum (in lateral view) with more steeply declined frontal margin. Additionally, D. evritaniensis , D. etoliensis and D. zbuzeki sp. nov. have the semi-erect pubescence along the whole length of the pronotal sides ( Figs 125–127 , 162 ). Dima hladilorum has a more convex scutellum and more robust apical lobe of paramera ( Fig. 190 ) and D. zbuzeki sp. nov. is smaller and has distinctly longer apical lobe of paramera ( Fig. 221 ).

Description. Holotype, male. Body medium-sized, 12.8 mm long, 5.1 mm wide, matt, covered with very short dense pubescence. Body dark brown, with paler, reddish-brown antennae, legs, elytral outer margins and suture; pubescence yellowish ( Fig. 92 ).

Head including eyes 0.6 times as wide as pronotum, frons with shallow depression; punctation dense and coarse, intervals between punctures dull; pubescence on head semi-erect, more erect in anterior part of frons, directed forwards. Supraantennal carina weakly defined. Antennae moderately long, surpassing the posterior angles of pronotum by more than three segments; antennomeres II–III subtriangular, about 1.5 times longer than wide, II slightly longer than III, together 1.2 times longer than antennomere IV, length ratio of antennomeres II–IV 1.1: 1.0: 1.9, antennomeres IV–X gradually widened apically, slightly more than 2 times longer than wide, apical antennomere subacute apically; surface of antennomeres covered with moderately long, semi-erect setae.

Pronotum 1.4 times wider than long, widest at middle, moderately convex dorsally; sides evenly convex, gradually slightly curved anteriorly in lateral view; anterior angles obtuse; posterior angles prominent, sharp, distinctly produced outward; punctation dense, fine, shallow, punctures on disc equally distributed, of about the same size; intervals between punctures shorter or of the same length as a diameter of puncture, shiny, more flattened posteriorly. Pubescence on pronotal disc very short, decumbent; that on sides decumbent at the anterior two thirds and semi-erect at the posterior third, longer setae at hind angles ( Fig. 152 ). Prosternum finely punctate, with short decumbent to semi-erect setae; prosternal lobe moderately coarsely punctate, with longer pubescence.

Scutellum heart-shaped, flattened, in lateral view slightly raised above the plane of elytra, widest at two thirds; frontal margin rounded in lateral view, gradually declined; apex widely rounded, punctation fine, dense; pubescence fine, moderately long, semi-erect. Elytra sub-parallel, 3.3 times longer than pronotum, moderately convex, widest at middle, from middle gradually narrowing towards apex; striae well developed along the whole elytral length; surface of interstices flattened, moderately shiny, densely and finely punctate; punctures suboval, equally distributed, of the same size; pubescence short, dense.

Aedeagus elongate; paramera long, narrow, its apical lobe robust, subapical tooth short, blunt ( Fig. 209 ).

Female ( Figs 93–95 ). Like male, but with suboval body, shorter antennae (surpassing the posterior angles of pronotum about two segments), head 0.5 times as wide as pronotum, pronotum wider than in male (1.5 times wider than long), scutellum wider. The sclerotized spines of bursa copulatrix as in Fig. 238 .

Intraspecific variability. Some specimens have rugose, more convex and more densely punctate pronotum ( Fig. 94 ), similarly as in some other Dima species (e.g. D. fthiotidensis , D. scutellaris ). Body coloration is variable; some specimens have paler, reddish-brown some parts of body, e.g. antennae, pronotum, legs.

Distribution. Greece (Thesally: Mt. Pelion; Fig. 245 ).

Etymology. The name " pelionensis " refers to Mt. Pelion, Greece, the type locality of the new species.

Remark. Schimmel & Platia (2008) described D. fokidensis with the holotype from Mt. Iti and paratypes from Mt. Pelion. That holotype is conspecific with D. fthiotidensis and the paratypes are here assigned to D. pelionensis sp. nov. ( Fig. 95 ).

Figures

FIGURE 245. The distribution of Dima species in the Central Greece.

FIGURES 90 – 104. Habitus images of Dima species. Dima pelikani sp. nov., 90 — paratype, female, 91 — paratype, male; D. pelionensis sp. nov., 92 — holotype, male, 93 — paratype, female, 94 — paratype, female; 95 — paratype, female, paratype of D. fokidensis; D. raineri Wurst, 1997, 96 — holotype, male, 97 — non-type specimen, male, 98 — non-type specimen, female, 99 — holotype of D. pindosensis Schimmel & Platia, 2008, male, 100 — male paratype of D. macedonica from Mt. Tomorr, Albania; 101 — Dima sp. from Librazhd, Albania; D. schimmeli Platia & Gudenzi, 2009, 102 — holotype, male, 103 — non-type specimen, male, 104 — non-type specimen, female. Not to scale.

FIGURES 143 – 154. Pronotal details of Dima species. 143 — D. macedonica Schimmel, 1993, male; 144 — D. marvani Mertlik & Dusanek, 2006, holotype, male; 145 — D. moraveci Schimmel & Platia, 2008, holotype, female; 146 — D. neumanni Platia, 2013, female; 147 — D. olympica Meschnigg, 1934, male; 148 — D. orientalis sp. nov., holotype, male; 149 — D. parnonensis Schimmel & Platia, 2008, female; 150 — D. pecoudi Fleutiaux, 1944, male; 151 — D. pelikani sp. nov., holotype, male; 152 — D. pelionensis sp. nov., paratype, female, 153 — D. pelionensis sp. nov., paratype, female paratype of D. fokidensis; 154 — D. raineri Wurst, 1997, female. Not to scale.

FIGURES 202 – 211. Aedeagus images of Dima species. 202 — D. olympica Meschnigg, 1934, non-type specimen; 203 — D. orientalis sp. nov., holotype; 204 — D. parnonensis Schimmel & Platia 2008, non-type specimen; 205 — D. parnonensis, nontype specimen; 206 — D. pecoudi Fleutiaux, 1944, paratype of D. evritaniensis; 207 — D. pecoudi, non-type specimen; 208 — D. pelikani sp. nov., holotype; 209 — D. pelionensis sp. nov., paratype; 210 — D. raineri Wurst, 1997, holotype; 211 — D. raineri, non-type specimen. Not to scale.

FIGURES 234 – 242. The sclerotized spines of bursa copulatrix of Dima species. 234 — D. olympica Meschnigg, 1934, nontype specimen; 235 — D. orientalis sp. nov., paratype; 236 — Dima sp. from Xanthi, Greece; 237 — D. pelikani sp. nov., paratype; 238 — D. pelionensis sp. nov., paratype of D. fokidensis Schimmel & Platia, 2008; 239 — D. schimmeli Platia & Gudenzi, 2009, non-type specimen; 240 — D. schnitteri Platia, 2013, paratype; 241 — D. vonickai sp. nov., paratype; 242 — D. zbuzeki sp. nov, paratype. Not to scale.

FIGURES 15 – 29. Habitus images of Dima species. Dima dalmatina Küster, 1844, 15 — syntype, male, 16 — non-type male from Vosac, Croatia, 17 — non-type female from Vosac, Croatia, 18 — non-type male from Rumija Mts., Montenegro, 19 — nontype female from Rumija Mts., Montenegro; D. dusaneki sp. nov., 20 — holotype, male, 21 — paratype, female; D. elateroides Charpentier, 1825, 22 — non-type male from Serbia; 23 — non-type male from Slovenia, 24 — non-type female from Croatia; 25 — D. etoliensis Platia, 2012, holotype, female; D. evritaniensis Schimmel & Platia, 2008, 26 — paratype, female, Panetoliko Mts., 27 — non-type male from Timfristos Mts., 28 — non-type female from Timfristos Mts.; 29 — D. fialai sp. nov., holotype, male. Not to scale.

FIGURES 30 – 44. Habitus images of Dima species. 30 — D. fialai sp. nov., paratype, female; D. florinensis Platia, 2012, 31 — holotype, female, 32 — non-type male from Varnous Mts., 33 — non-type female from Varnous Mts., 34 — non-type female from Galičica Mts.; D. fthiotidensis Schimmel & Platia, 2008, 35 — paratype, female, 36 — holotype of D. fokidensis Schimmel & Platia, 2008, female, 37 — holotype of D. scutellaris Platia, 2010, female, 38 — non-type male from Kallidromo Mts., 39 — nontype male from Vardoussia Mts., 40 — non-type female from Vardoussia Mts., 41 — non-type female from Vardoussia Mts.; D. giachinoi Schimmel & Platia 2008, 42 — paratype, female, 43 — non-type specimen, male, 44 — non-type specimen, female. Not to scale.

FIGURES 119 – 130. Pronotal details of Dima species. 119 — D. arndti Platia, 2013, male; 120 — D. assingi Schimmel & Platia, 2008, female; 121 — D. bruhai sp. nov., holotype, male; 122 — D. dalmatina Küster, 1844, male; 123 — D. dusaneki sp. nov., holotype, male; 124 — D. elateroides Charpentier, 1825, male; 125 — D. etoliensis Platia, 2012, holotype, female; 126 — D. evritaniensis Schimmel & Platia, 2008, paratype, female; 127 — D. evritaniensis, male; 128 — D. fialai sp. nov., holotype, male; 129 — D. florinensis Platia, 2012, female; 130 — D. fthiotidensis Schimmel & Platia, 2008, male. Not to scale.

FIGURES 131 – 142. Pronotal details of Dima species. 131 — D. fthiotidensis Schimmel & Platia, 2008, paratype, female; 132 — D. scutellaris Platia, 2010, holotype, female; 133 — D. fthiotidensis, male; 134 — D. fthiotidensis, holotype of D. fokidensis Schimmel & Platia, 2008; 135 — D. giachinoi Schimmel & Platia 2008, female; 136 — D. hirtipennis Platia, 2011, holotype, female; 137 — D. hladilorum Schimmel, 1987, male; 138 — D. kozufensis sp. nov., paratype, female; 139 — Dima lebenbaueri Schimmel & Platia 2008, holotype, male; 140 — Dima lebenbaueri, non-type specimen, female; 141 — D. macedonica Schimmel, 1993, holotype, female, 142 — D. macedonica, female paratype of D. hladilorum from Galičica Mts. Not to scale.

FIGURES 155 – 162. Pronotal details of Dima species. 155 — D. raineri Wurst, 1997, holotype, male; 156 — D. schimmeli Platia & Gudenzi, 2009, female; 157 — D. schnitteri Platia, 2013, female; 158 — D. vailatii Schimmel & Platia 2008, holotype, male; 159 — D. vailatii, male; 160 — D. riesei Platia, 2010, holotype, male; 161 — D. vonickai sp. nov., holotype, male; 162 — D. zbuzeki sp. nov., holotype, male. Not to scale.

FIGURES 182 – 191. Aedeagus images of Dima species. 182 — D. elateroides Charpentier, 1825, non-type specimen from Serbia; 183 — D. evritaniensis Schimmel & Platia, 2008, non-type specimen from Timfristos Mts.; 184 — D. fialai sp. nov., holotype; 185 — D. florinensis Platia, 2012, non-type specimen; 186 — D. fthiotidensis Schimmel & Platia, 2008, non-type specimen from Kallidromo Mts.; 187 — D. fthiotidensis, non-type specimen from Vardoussia Mts.; 188 — D. giachinoi Schimmel & Platia 2008, non-type specimen; 189 — Dima sp. from Mani; 190 — D. hladilorum Schimmel, 1987, non-type specimen; 191 — D. kozufensis, sp. nov., holotype. Not to scale.

FIGURES 212 – 221. Aedeagus images of Dima species. 212 — D. raineri Wurst, 1997, paratype of D. macedonica; 213 — D. raineri, holotype of D. pindosensis Schimmel & Platia, 2008; 214 — D. schimmeli Platia & Gudenzi, 2009, holotype; 215 — D. schimmeli, non-type specimen; 216 — D. schnitteri Platia, 2013, holotype; 217 — D. vailatii Schimmel & Platia, 2008, holotype; 218 — D. vailatii, non-type specimen; 219 — D. vailatii, holotype of D. riesei Platia, 2010; 220 — D. vonickai sp. nov., holotype; 221 — D. zbuzeki sp. nov., holotype. Not to scale.