Dima macedonica Schimmel, 1993

Mertlik, Josef, Németh, Tamás & Kundrata, Robin, 2017, Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula, Zootaxa 4220 (1), pp. 1-63 : 22

Dima macedonica Schimmel, 1993

Figs 57–61 , 141–143 , 164, 168 , 194–197 , 244 .

Dima macedonica Schimmel, 1993: 247 ; Schimmel 1996: 145; Wurst 1997: 3; Cate 2007: 184; Schimmel & Platia 2008: 569.

Type depositories. Holotype, ♂ (PCPC), 2 paratypes, ♂♂ (PCPC, PCRS).

Type locality. Macedonia, Galičica , 1600 m. [YU: Makedonija].

Redetermined material. A male paratype of D. macedonica from Albania: Mt. Tomorr ( PCGP) was redetermined as D. raineri ( Fig. 100 ). Some specimens recorded under D. macedonica in Schimmel & Platia (2008) were redetermined as D. elateroides ( Croatia, Montenegro; 6 ex.) and D. raineri ( Greece; 7 ex.). For a female specimen from Librazhd, Albania ( Fig. 101 ), identified as D. macedonica by Schimmel & Platia (2008), see the remarks under D. raineri . Three paratypes of D. hladilorum from Macedonia (Galičica Mts) belong under D. macedonica ( Fig. 61 ).

New material. MACEDONIA: Galičica Mts., Livada pass, Mt. Magaro , 1650–1800 m, 9.–10. VI.2008, 1 ♀, P. Vonička leg. ( PCJM) ; dtto, 5 ♂♂, 3 ♀♀, L. Blažej leg. (PCPB); NP Galičica – Livada, Trpejca env., 1401 m (40°57'48"N, 20°48'49"E), 2.–3. VI.2011, 1 ♂, D. Navrátil leg. ( PCJM) ; distr. Ohrid, Galičica plateau, Trpejca env., beechwood, 1540 m (40°57'42.18"N, 20°48'55.76"E), 24. V.2014, 4 ♂♂, 3 ♀♀, J. Mertlik leg. ( PCJM) ; dtto, 1 ♂, 1 ♀, B. Zbuzek leg. (PCBZ); dtto, 7 ♂♂, 1 ♀, P. Brůha leg. (PCPB); distr. Ohrid, Galičica plateau, beechwood, near pass, 1590 m (40°57'7.72"N, 20°49'56.16"E), 24. V.2014, 28 ♂♂, 4 ♀♀, J. Mertlik leg. ( PCJM) ; dtto, 19 ♂♂, B. Zbuzek leg. (PCBZ); dtto, 22 ♂♂, 2 ♀♀, P. Brůha leg. (PCPB); Ohrid, PN Galičica, 2 km S of Lako Signoj , 1940 m, 40°56.285'N, 20°49.407'E, 7. VI.2015, 3 ♀♀, Richard Škoda leg. ( PCPB) ; Pelagonia reg., 1.5 km SW Ehloec (41°27'3.88"N, 20°44'30.19"E), 6.VI.2014, Fagus forest, swept & beaten, 3 ♂♂, leg. Dusanek, Kundrata & Németh ( PCRK) ; dtto, 1 ♀ (HNHM); dtto, 4 ♂♂, 4 ♀♀, V. Dušánek leg. (PCVD).

Diagnosis. Dima macedonica is a medium-sized species (body length: 9.8–13.2 mm) which belongs to the group of species with more convex, suboval-shaped body and shiny elytral intervals with very sparsely distributed thin setae, similarly as e.g. D. florinensis and D. raineri ( Figs 57–61 ). Dima macedonica is characterized by the scutellum raised above the plane of the elytra (clearly visible in lateral view; Fig. 164 ), distinctly double-sized punctation of pronotum, and the pubescence of the pronotal sides which is semi-erect along the whole margin length but in anterior half about 0.5 times as long as in posterior half ( Figs 141–143 ). All these characters can be used for the separation of D. macedonica and D. raineri , which has the scutellum on the same plane as the elytra, double punctation of pronotum less distinct, with punctures more similar in size to each other ( Figs 154–155 , 165 ), and pubescence on the anterior halfs of pronotal sides shorter and decumbent ( Figs 154–155 ). Dima florinensis shares with D. macedonica the double punctation of pronotum, but differs in the scutellum, which is on the same plane as the elytra, longer pubescence of pronotal sides, which is of about the same length along the whole margin length ( Fig. 129 ), and by the shorter apical lobe of paramera ( Figs 185 , 194–197 ).

Intraspecific variability. The shape of antennomeres II and III is variable. Most specimens of D. macedonica have antennomeres II and III subtriangular, but in some specimens they are more elongate, as in most specimens of D. raineri (see Wurst 1997). Similar variability occurs also in D. raineri , when some specimens have similar antennomeres II and III as the typical representatives of D. macedonica . Specimens of the northern population of D. macedonica (near Ehloec) have more convex base of scutellum.

Distribution. Macedonia (Galičica Mts., Ehloec; Fig. 244 ).

Figures

FIGURE 244. The distribution of Dima species in Albania, Bulgaria, Macedonia and in the northern Greece.

FIGURES 45 – 59. Habitus images of Dima species. 45 — Dima hirtipennis Platia, 2011, holotype, female, 46 — Dima sp. from Mani; D. hladilorum Schimmel, 1987, 47 — paratype, female, 48 — non-type specimen, male, 49 — non-type specimen, female; D. kozufensis sp. nov., 50 — holotype, male, 51 — paratype, female, 52 — paratype, female, paratype of D. lebenbaueri Schimmel & Platia, 2008; D. lebenbaueri Schimmel & Platia 2008, 53 — holotype, male, 54 — paratype, female, 55 — non-type specimen, male, 56 — non-type specimen, female; D. macedonica Schimmel, 1993, 57 — holotype, male, 58 — non-type male from Galičica, 59 — non-type female from Galičica Mts. Not to scale.

FIGURES 60 – 74. Habitus images of Dima species. Dima macedonica Schimmel, 1993, 60 — non-type male from Ehloec, 61 — female paratype of D. hladilorum Schimmel, 1987 from Galičica Mts.; D. marvani Mertlik & Dusanek, 2006, 62 — holotype, male, 63 — female paratype from Lovčen Mts., 64 — female paratype from Rumija Mts., 65 — non-type male from Rumija Mts.; 66 — D. moraveci Schimmel & Platia 2008, holotype, female; D. neumanni Platia, 2013, 67 — holotype, male, 68 — non-type male from Erimanthos Mts., 69 — non-type female from Erimanthos Mts., 70 — non-type male from Aroania Mts., 71 — non-type female from Aroania Mts., 72 — non-type female from Aroania Mts., 73 — non-type female from Panachaiko Mts., 74 — non-type female from Panachaiko Mts. Not to scale.

FIGURES 131 – 142. Pronotal details of Dima species. 131 — D. fthiotidensis Schimmel & Platia, 2008, paratype, female; 132 — D. scutellaris Platia, 2010, holotype, female; 133 — D. fthiotidensis, male; 134 — D. fthiotidensis, holotype of D. fokidensis Schimmel & Platia, 2008; 135 — D. giachinoi Schimmel & Platia 2008, female; 136 — D. hirtipennis Platia, 2011, holotype, female; 137 — D. hladilorum Schimmel, 1987, male; 138 — D. kozufensis sp. nov., paratype, female; 139 — Dima lebenbaueri Schimmel & Platia 2008, holotype, male; 140 — Dima lebenbaueri, non-type specimen, female; 141 — D. macedonica Schimmel, 1993, holotype, female, 142 — D. macedonica, female paratype of D. hladilorum from Galičica Mts. Not to scale.

FIGURES 143 – 154. Pronotal details of Dima species. 143 — D. macedonica Schimmel, 1993, male; 144 — D. marvani Mertlik & Dusanek, 2006, holotype, male; 145 — D. moraveci Schimmel & Platia, 2008, holotype, female; 146 — D. neumanni Platia, 2013, female; 147 — D. olympica Meschnigg, 1934, male; 148 — D. orientalis sp. nov., holotype, male; 149 — D. parnonensis Schimmel & Platia, 2008, female; 150 — D. pecoudi Fleutiaux, 1944, male; 151 — D. pelikani sp. nov., holotype, male; 152 — D. pelionensis sp. nov., paratype, female, 153 — D. pelionensis sp. nov., paratype, female paratype of D. fokidensis; 154 — D. raineri Wurst, 1997, female. Not to scale.

FIGURES 163 – 171. Scutellum-shape of Dima species. 163 — D. bruhai sp. nov.; 164 — D. macedonica Schimmel, 1993; 165 — D. raineri Wurst, 1997. Elytral details of Dima species. 166 — D. fthiotidensis Schimmel & Platia, 2008; 167 — D. hirtipennis Platia, 2011; 168 — D. macedonica Schimmel, 1993; 169 — D. olympica Meischnigg, 1934; 170 — D. parnonensis Schimmel & Platia, 2008; 171 — D. pecoudi Fleutiaux, 1943. Not to scale.

FIGURES 192 – 201. Aedeagus images of Dima species. 192 — D. lebenbaueri Schimmel & Platia 2008, holotype; 193 — D. lebenbaueri, non-type specimen; 194 — D. macedonica Schimmel, 1993, holotype; 195 — D. macedonica, paratype; 196 — D. macedonica, non-type specimen from Galičica Mts.; 197 — D. macedonica, non-type specimen from Ehloec; 198 — D. marvani Mertlik & Dusanek, 2006, holotype; 199 — D. neumanni Platia, 2013, holotype; 200 — D. neumanni, non-type specimen from Erimanthos Mts.; 201 — D. neumanni, non-type specimen from Aroania Mts. Not to scale.

FIGURES 90 – 104. Habitus images of Dima species. Dima pelikani sp. nov., 90 — paratype, female, 91 — paratype, male; D. pelionensis sp. nov., 92 — holotype, male, 93 — paratype, female, 94 — paratype, female; 95 — paratype, female, paratype of D. fokidensis; D. raineri Wurst, 1997, 96 — holotype, male, 97 — non-type specimen, male, 98 — non-type specimen, female, 99 — holotype of D. pindosensis Schimmel & Platia, 2008, male, 100 — male paratype of D. macedonica from Mt. Tomorr, Albania; 101 — Dima sp. from Librazhd, Albania; D. schimmeli Platia & Gudenzi, 2009, 102 — holotype, male, 103 — non-type specimen, male, 104 — non-type specimen, female. Not to scale.

FIGURES 155 – 162. Pronotal details of Dima species. 155 — D. raineri Wurst, 1997, holotype, male; 156 — D. schimmeli Platia & Gudenzi, 2009, female; 157 — D. schnitteri Platia, 2013, female; 158 — D. vailatii Schimmel & Platia 2008, holotype, male; 159 — D. vailatii, male; 160 — D. riesei Platia, 2010, holotype, male; 161 — D. vonickai sp. nov., holotype, male; 162 — D. zbuzeki sp. nov., holotype, male. Not to scale.

FIGURES 119 – 130. Pronotal details of Dima species. 119 — D. arndti Platia, 2013, male; 120 — D. assingi Schimmel & Platia, 2008, female; 121 — D. bruhai sp. nov., holotype, male; 122 — D. dalmatina Küster, 1844, male; 123 — D. dusaneki sp. nov., holotype, male; 124 — D. elateroides Charpentier, 1825, male; 125 — D. etoliensis Platia, 2012, holotype, female; 126 — D. evritaniensis Schimmel & Platia, 2008, paratype, female; 127 — D. evritaniensis, male; 128 — D. fialai sp. nov., holotype, male; 129 — D. florinensis Platia, 2012, female; 130 — D. fthiotidensis Schimmel & Platia, 2008, male. Not to scale.

FIGURES 182 – 191. Aedeagus images of Dima species. 182 — D. elateroides Charpentier, 1825, non-type specimen from Serbia; 183 — D. evritaniensis Schimmel & Platia, 2008, non-type specimen from Timfristos Mts.; 184 — D. fialai sp. nov., holotype; 185 — D. florinensis Platia, 2012, non-type specimen; 186 — D. fthiotidensis Schimmel & Platia, 2008, non-type specimen from Kallidromo Mts.; 187 — D. fthiotidensis, non-type specimen from Vardoussia Mts.; 188 — D. giachinoi Schimmel & Platia 2008, non-type specimen; 189 — Dima sp. from Mani; 190 — D. hladilorum Schimmel, 1987, non-type specimen; 191 — D. kozufensis, sp. nov., holotype. Not to scale.