Dima vonickai, Mertlik, Josef, Németh, Tamás & Kundrata, Robin, 2017

Mertlik, Josef, Németh, Tamás & Kundrata, Robin, 2017, Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula, Zootaxa 4220 (1), pp. 1-63 : 33-34

Dima vonickai sp. nov.

Figs 114–115 , 161 , 220 , 241 , 244 .

Type material. Holotype: ALBANIA: Zepës Mts. , Lar env., 1600 m, 8.VI.2014, ♂, P. Moravec leg. ( PCJM) . Paratypes: ALBANIA: Zepës Mts. , Lar env., 1600 m, 8.VI.2014, 1 ♀, P. Moravec leg. ( PCJM) ; dtto, 1 ♂, 1 ♀, P. Vonička, P. Moravec et L. Blažej leg. (PCPB).

Diagnosis. Dima vonickai sp. nov. belongs to the group of species with more convex, suboval-shaped body and shiny elytral intervals with very sparsely distributed thin setae ( Fig. 114–115 ). This species is like D. fialai sp. nov., D. dusaneki sp. nov. and D. raineri based on the pubescence of the pronotal lateral margin, which is short, almost decumbent in anterior part and longer, semi-erect posteriorly ( Fig. 161 ). Dima fialai sp. nov. and D. dusaneki sp. nov. differ from D. vonickai sp. nov. by more robust body, wide and convex pronotum ( Figs 20–21, 29–30 ), and shorter apical lobe of paramera ( Figs 181 , 184 ). Dima raineri has pronotal punctures of two different sizes (almost the same size in D. vonickai sp. nov.; Figs 154–155, 161 ) and differently shaped paramera ( Figs 210– 213, 220 ).

Description. Holotype, male. Body medium-sized, 12.5 mm long, 5.1 mm wide, shiny, covered with short dense pubescence on pronotum and sparse on elytra. Body black, with paler, reddish-brown legs, antennae, elytral sutura and margins; pubescence yellowish ( Fig. 114 ).

Head including eyes 0.6 times as wide as pronotum, frons with shallow v-shaped depression; punctation sparse and coarse, intervals between punctures shiny; pubescence on head semi-erect, more erect in anterior part of frons, directed forwards. Supraantennal carina weakly developed, short. Antennae moderately long, surpassing the posterior angles of pronotum of about 3.5 segments; antennomeres II and III subtriangular, more than 1.5 times longer than wide, subequal, together about 1.4 times longer than antennomere IV, length ratio of antennomeres II– IV 1.0: 1.0: 1.4, antennomeres IV to X gradually widened apically, less than 2 times longer than wide, apical antennomere longest, subacute apically; surface of antennomeres covered with moderately long, semi-erect setae.

Pronotum 1.4 times wider than long, widest just behind middle, moderately convex dorsally; sides convex; anterior angles obtuse; posterior angles prominent, sharp, slightly produced outward; punctation sparse, moderately fine, punctures on disc equally distributed, of about the same size; intervals between punctures of 2–3 times a diameter of puncture, flat, shiny. Pubescence on pronotal disc moderately dense, short, decumbent; that on pronotal sides in anterior two thirds decumbent, in posterior third semi-erect; setae in hind angles long, erect ( Fig. 161 ). Prosternum finely, sparsely punctate, interstices glabrous, with short decumbent to semi-erect setae, denser at margins; prosternal lobe moderately coarsely punctate, with longer semi-erect pubescence.

Scutellum sub-pentagonal, with sides sub-parallel, in lateral view slightly raised above the plane of elytra, widest at two thirds; frontal margin rounded, gradually declined, apex rounded; punctation fine, dense; pubescence fine, short, semi-erect. Elytra suboval, 3.3 times longer than pronotum, convex, widest at posterior two thirds; striae well developed along the whole elytral length, some of them irregularly interrupted by coarse sparse punctures; surface of interstices flattened, shiny, sparsely punctate; punctures shallow, suboval, equally distributed, of the same size; pubescence short, very sparse.

Aedeagus elongate; paramera long, narrow, its apical lobe elongate, subapical tooth subacute ( Fig. 220 ).

Female ( Fig. 115 ). Very like male (in contrast with most other Dima spp.), but more robust, with shorter antennae (surpassing the posterior angles of pronotum about two segments) and slightly wider scutellum. The sclerotized spines of bursa copulatrix as in Fig. 241 .

Intraspecific variability. Body length: 10.7–12.3 mm.

Distribution. Albania (Zepës Mts.; Fig. 244 ).

Etymology. The specific epithet is a patronym in honor of Mr. Pavel Vonička (Liberec, Czech Republic), who collected a part of the type series.

Figures

FIGURE 244. The distribution of Dima species in Albania, Bulgaria, Macedonia and in the northern Greece.

FIGURES 105 – 118. Habitus images of Dima species. Dima schnitteri Platia, 2013, 105 — holotype, male, 106 — paratype, female, 107 — non-type specimen, male, 108 — non-type specimen, female; D. vailatii Schimmel & Platia 2008, 109 — holotype, male, 110 — paratype, female, 111 — non-type specimen, male, 112 — non-type specimen, female, 113 — holotype of D. riesei Platia, 2010, male; D. vonickai sp. nov., 114 — holotype, male, 115 — paratype, female; D. zbuzeki sp. nov., 116 — holotype, male, 117 — paratype, female, 118 — paratype, male. Not to scale.

FIGURES 155 – 162. Pronotal details of Dima species. 155 — D. raineri Wurst, 1997, holotype, male; 156 — D. schimmeli Platia & Gudenzi, 2009, female; 157 — D. schnitteri Platia, 2013, female; 158 — D. vailatii Schimmel & Platia 2008, holotype, male; 159 — D. vailatii, male; 160 — D. riesei Platia, 2010, holotype, male; 161 — D. vonickai sp. nov., holotype, male; 162 — D. zbuzeki sp. nov., holotype, male. Not to scale.

FIGURES 212 – 221. Aedeagus images of Dima species. 212 — D. raineri Wurst, 1997, paratype of D. macedonica; 213 — D. raineri, holotype of D. pindosensis Schimmel & Platia, 2008; 214 — D. schimmeli Platia & Gudenzi, 2009, holotype; 215 — D. schimmeli, non-type specimen; 216 — D. schnitteri Platia, 2013, holotype; 217 — D. vailatii Schimmel & Platia, 2008, holotype; 218 — D. vailatii, non-type specimen; 219 — D. vailatii, holotype of D. riesei Platia, 2010; 220 — D. vonickai sp. nov., holotype; 221 — D. zbuzeki sp. nov., holotype. Not to scale.

FIGURES 234 – 242. The sclerotized spines of bursa copulatrix of Dima species. 234 — D. olympica Meschnigg, 1934, nontype specimen; 235 — D. orientalis sp. nov., paratype; 236 — Dima sp. from Xanthi, Greece; 237 — D. pelikani sp. nov., paratype; 238 — D. pelionensis sp. nov., paratype of D. fokidensis Schimmel & Platia, 2008; 239 — D. schimmeli Platia & Gudenzi, 2009, non-type specimen; 240 — D. schnitteri Platia, 2013, paratype; 241 — D. vonickai sp. nov., paratype; 242 — D. zbuzeki sp. nov, paratype. Not to scale.

FIGURES 15 – 29. Habitus images of Dima species. Dima dalmatina Küster, 1844, 15 — syntype, male, 16 — non-type male from Vosac, Croatia, 17 — non-type female from Vosac, Croatia, 18 — non-type male from Rumija Mts., Montenegro, 19 — nontype female from Rumija Mts., Montenegro; D. dusaneki sp. nov., 20 — holotype, male, 21 — paratype, female; D. elateroides Charpentier, 1825, 22 — non-type male from Serbia; 23 — non-type male from Slovenia, 24 — non-type female from Croatia; 25 — D. etoliensis Platia, 2012, holotype, female; D. evritaniensis Schimmel & Platia, 2008, 26 — paratype, female, Panetoliko Mts., 27 — non-type male from Timfristos Mts., 28 — non-type female from Timfristos Mts.; 29 — D. fialai sp. nov., holotype, male. Not to scale.

FIGURES 30 – 44. Habitus images of Dima species. 30 — D. fialai sp. nov., paratype, female; D. florinensis Platia, 2012, 31 — holotype, female, 32 — non-type male from Varnous Mts., 33 — non-type female from Varnous Mts., 34 — non-type female from Galičica Mts.; D. fthiotidensis Schimmel & Platia, 2008, 35 — paratype, female, 36 — holotype of D. fokidensis Schimmel & Platia, 2008, female, 37 — holotype of D. scutellaris Platia, 2010, female, 38 — non-type male from Kallidromo Mts., 39 — nontype male from Vardoussia Mts., 40 — non-type female from Vardoussia Mts., 41 — non-type female from Vardoussia Mts.; D. giachinoi Schimmel & Platia 2008, 42 — paratype, female, 43 — non-type specimen, male, 44 — non-type specimen, female. Not to scale.

FIGURES 172 – 181. Aedeagus images of Dima species. 172 — D. arndti Platia, 2013, holotype; 173 — D. arndti, non-type specimen; 174 — Dima sp. from Crete; 175 — D. assingi Schimmel & Platia, 2008, holotype; 176 — D. assingi, non-type specimen from Kilini Mts.; 177 — D. peloponnensis Schimmel & Platia 2008, holotype; 178 — D. bruhai sp. nov., holotype; 179 — D. dalmatina Küster, 1844, non-type specimen from Sutorman, Montenegro; 180 — D. dalmatina, non-type specimen from Vosec, Croatia; 181 — D. dusaneki sp. nov., holotype. Not to scale.

FIGURES 182 – 191. Aedeagus images of Dima species. 182 — D. elateroides Charpentier, 1825, non-type specimen from Serbia; 183 — D. evritaniensis Schimmel & Platia, 2008, non-type specimen from Timfristos Mts.; 184 — D. fialai sp. nov., holotype; 185 — D. florinensis Platia, 2012, non-type specimen; 186 — D. fthiotidensis Schimmel & Platia, 2008, non-type specimen from Kallidromo Mts.; 187 — D. fthiotidensis, non-type specimen from Vardoussia Mts.; 188 — D. giachinoi Schimmel & Platia 2008, non-type specimen; 189 — Dima sp. from Mani; 190 — D. hladilorum Schimmel, 1987, non-type specimen; 191 — D. kozufensis, sp. nov., holotype. Not to scale.

FIGURES 143 – 154. Pronotal details of Dima species. 143 — D. macedonica Schimmel, 1993, male; 144 — D. marvani Mertlik & Dusanek, 2006, holotype, male; 145 — D. moraveci Schimmel & Platia, 2008, holotype, female; 146 — D. neumanni Platia, 2013, female; 147 — D. olympica Meschnigg, 1934, male; 148 — D. orientalis sp. nov., holotype, male; 149 — D. parnonensis Schimmel & Platia, 2008, female; 150 — D. pecoudi Fleutiaux, 1944, male; 151 — D. pelikani sp. nov., holotype, male; 152 — D. pelionensis sp. nov., paratype, female, 153 — D. pelionensis sp. nov., paratype, female paratype of D. fokidensis; 154 — D. raineri Wurst, 1997, female. Not to scale.

FIGURES 202 – 211. Aedeagus images of Dima species. 202 — D. olympica Meschnigg, 1934, non-type specimen; 203 — D. orientalis sp. nov., holotype; 204 — D. parnonensis Schimmel & Platia 2008, non-type specimen; 205 — D. parnonensis, nontype specimen; 206 — D. pecoudi Fleutiaux, 1944, paratype of D. evritaniensis; 207 — D. pecoudi, non-type specimen; 208 — D. pelikani sp. nov., holotype; 209 — D. pelionensis sp. nov., paratype; 210 — D. raineri Wurst, 1997, holotype; 211 — D. raineri, non-type specimen. Not to scale.