Eratomyia, Published, 2007

DE SOUZA AMORIM, DALTON & RINDAL, EIRIK, 2007, Phylogeny of the Mycetophiliformia, with proposal of the subfamilies Heterotrichinae, Ohakuneinae, and Chiletrichinae for the Rangomaramidae (Diptera, Bibionomorpha), Zootaxa 1535 (1), pp. 1-92 : 14

Eratomyia , n. gen.

( Figs. 7–15 )

Type species. Eratomyia magnifica sp. n., here designated

Diagnosis. First sector of m-cu longer than in other genera of subfamily, m-cu reaching CuA more distally. M 1+2 /M 3+4 shorter than m-cu, r-m long. Tergites 1-4 and 6 shining brown, tergite 5 yellow. Midpleural pit absent.

Etymology. The species name is feminine, from the Greek eratos, lovely, meaning a lovely fly.

Comments. This genus, which is known only from the type species, from Ecuador, appears in the phylogenetic analysis (below) as the sister group of Chiletricha , which is known from Chile, southern Argentina, and southern Brazil. It differs from the Chiletricha species in wing features, with a much wider basal cell, R 5 separated from C, and M 4 originating from CuA not as basally in the wing ( Figs. 10 , 16 ). Mouthparts are fairly similar ( Figs. 7 , 17 ). In the thorax, the mesepimeron of Eratomyia is more slender on its ventral half, whereas the laterotergite is slightly larger ( Figs. 8 , 18 ). Tergite 9 has strong spines on a pair of distal projections ( Fig. 12–15 ), as also seen in Chiletricha ( Fig. 19 ), Rhynchoheterotricha , and Insulatricha , and the gonocoxites have a pair of unique digitiform projections bearing setae and a pair of spines.

Figures

FIGURES 7–10. Eratomyia magnifica, sp. n. 7. Head (apical maxillary palpal segments not represented), lateral view. 8. Thorax. 9. Front tibial apex, inner view. 10. Wing.

FIGURES 11–15. Eratomyia magnifica, sp. n. 11. Male terminalia, posterior view. 12. Male terminalia, lateral view. 13. Gonostyle, external view. 14. Aedeagus and gonocoxal bridge. 15. Tergites VIII–X and cerci, posterior view.

FIGURES 16–19. Chiletricha. 16, 17. Chiletricha seminuda (Freeman). 16. Wing. 17. Mouthparts, posterior view. 18, 19. Chiletricha marginata (Edwards). 18. Thorax. 19. Male terminalia, dorsal view.