Lyroda triloba Say, 1837

Khvir, Viktor I. & Pulawski, Wojciech J., 2020, A Revision of New World Lyroda Say, 1837 (Hymenoptera: Crabronidae), Proceedings of the California Academy of Sciences 66 (13), pp. 315-330 : 326

Lyroda triloba Say

Lyroda triloba Say, 1837:372 , sex not indicated. Holotype or syntypes: North America: no specific locality (lost). Neotype: ♀, Arkansas: Hampstead County: no specific locality (CAS), present designation. – Le Conte, 1859:755 (original description copied); Patton, 1880:387 (in checklist of North American Larrinae); Provancher, 1882:49 and 1883:631 (original description translated into French); Kohl, 1885:267 (in checklist of world Lyroda ); Cresson, 1887:277 (in catalog of North American Hymenoptera ); Ashmead, 1890:33 (in checklist of Hymenoptera of Colorado); W. Fox, 1894:534 (in revision of North American Larrinae); Dalla Torre, 1897:696 (in catalog of world Hymenoptera ); Ashmead, 1899:250 (in checklist of North American Crabronidae ); Bridwell, 1899:209 (Kansas: no specific locality); J. Smith, 1900:518 (in list of insects of New Jersey); H. Smith, 1908:374 (in revision of Nebraskan Sphecidae ); J. Smith, 1910:684 (in new list of insects of New Jersey: Camden County); F. Williams, 1914:174 (in revision of Larrinae of Kansas); Rohwer, 1916:683 (in catalog of Hymenoptera of Connecticut); Mickel, 1918:408 (in catalog of Nebraskan Sphecidae ); Rohwer in Viereck, 1925:683 (in key to Sphecidae of Connecticut; Branford); Iwata, 1933:7 (in list of world Lyroda ); Krombein in Muesebeck, Krombein and Townes, 1951:941 (in catalog of North American Hymenoptera ); R. Bohart and Menke, 1976:299 (in checklist of world Sphecidae ); Krombein, 1979:1633 (in catalog of North American Hymenoptera ). – As Tachytes trilobus : F. Smith, 1856:307 (new combination, in catalog of Hymenoptera in British Museum). – As Lyrops triloba : Cresson, 1862:238 (new combination, in catalog of North American Hymenoptera ). – As Larrada triloba : Cresson, 1873:213 (new combination, Texas).

Lyrops caliptera Say, 1837:373 . Nomen nudum or lapsus for Lyroda triloba . – Patton, 1880:387 (in checklist of North American Larrinae); Kohl, 1885:267 (in checklist of world Lyroda ).

RECOGNITION. Lyroda triloba differs from its New World congeners in having the propodeal side with well-defined ridges ( Fig. 19 ), the wing membrane dark, particularly in the female, and the large size (length of female 15.2–16.2 mm). In the other species, the propodeal side is either not ridged (most specimens) or (some L. subita ) has inconspicuous ridges, the wings are light colored (except dark in Floridian populations of L. subita ) and the female length does not exceed 13 mm. Unlike L. subita , the male of L. triloba has no median projection on the clypeal free margin (Fig. 18).

DESCRIPTION. Propodeal side with well-defined ridges ( Fig. 19 ), without spiracular groove. Tergum I in vast majority of specimens with several smaller ridges in addition to pair of main basomedian carinae. Body deep black. Wing membrane uniformly dark, particularly in female.

♀: Free margin of clypeus with three pairs of inconspicuous (almost invisible) teeth on each side of lamella ( Fig. 17 ). Clypeal setae dark. Pubescence of gastral terga all black. Length 15.2– 16.2 mm.

♂: Clypeal free margin inconspicuously, roundly arcuate mesally (Fig. 18). Clypeal setae silvery. Length 9.6–13.8 mm.

GEOGRAPHIC DISTRIBUTION (Fig. 20). Eastern North America, extending west beyond the 100th meridian only in South Dakota.

RECORDS. USA: Arkansas: Conway (1 ♀, CAS, 1 ♀, USNM), Hempstead (1 ♀, CAS, 2 ♀, USNM) . Georgia: Hiawassee (1 ♀, CNC) , Putnam Co. (5 ♀, USNM) . Kansas: no specific locality (1 ♂, CAS) . Louisiana: Tallulah (1 ♂, CAS) . Maryland: Beltsville (2 ♀, USNM) , Hudson (3 ♀, FSCA) . North Carolina: Stafford (1 ♂, CAS) , Washington Co. (1 ♀, FSCA) . South Dakota: Butte Co. (1 ♂, USNM) . Texas: McKinney (1 ♀, USNM) , no specific locality (1 ♀, USNM) . Virginia: Chance (1 ♀, USNM) , Norfolk (1 ♀, USNM) .

Figures

FIGURE 19. Lyroda triloba: propodeal side of female.

FIGURE 17. Lyroda triloba: female clypeus and mandibles