Glaresis costata Gordon and Hanley, 2014

Gordon, Robert D. & Hanley, Guy A., 2014, Systematic revision of American Glaresidae (Coleoptera: Scarabaeoidea), Insecta Mundi 2014 (333), pp. 1-91 : 10-12

Glaresis costata Gordon and Hanley , new species

Description. Male. Length 3.7 mm, width 1.9 mm; body form elongate, slightly widened from elytral base to apical 1/3 ( Fig. 4A ). Color yellowish brown. Head with clypeus and frons densely microreticulate, dull, with large, deep punctures, setae short, barely emergent from punctures; frons coarsely, densely microreticulate, feebly shiny, with few small, round tubercles, not setose; vertex with short, median basal carina, surface coarsely, densely microreticulate, feebly shiny, without tubercles or punctures. Clypeal apex truncate, smooth, narrow, angled posteriorly, lateral angles abruptly angulate ( Fig. 4B ). Mandible pair symmetrical; mesal tooth strong; lateral prominence strong, pronounced; outer margin angular. Pronotum with distinct central furrow extended from base nearly to apex, surface dull, coarsely, densely micoreticulate, with elongate, setae-bearing ridges arranged in various directions, setae distinct, slightly shorter than ridge ( Fig. 4C ). Anterior and lateral pronotal margins not bordered, anterior margin smooth medially, crenulate in lateral 1/4, lateral and basal margins entirely crenulate, basal margin bordered. Elytron with convex, somewhat carinate striae, surface feebly shiny, coarsely, densely microreticulate; each stria with small, closely spaced, nearly round, seta-bearing tubercle (presenting slightly carinate appearance), setae long, slightly longer than diameter of an interval puncture; interval with large, distinctly visible, rectangular punctures ( Fig. 4E ). Metasternum long, feebly shiny, coarsely, densely microreticulate throughout; entire surface with short, oblique, seta-bearing ridges; without trace of metasternal groove. Lateral protibial teeth unevenly spaced, basal tooth reduced. Mesotibia with 5 spines laterally, spinal bases widely spaced, outer apex of tibia not strongly projecting ( Fig. 4D ). Posterior margin of metatrochanter with 2 large teeth (usually an additional small tooth present) ( Fig. 4H ); posterosuperior surface without teeth. Metafemoral surface with widely scattered, elongate, setae-bearing ridges, finely microreticulate, dull throughout; shape slightly elongate; width to length ratio1.0:1.6, without flange on anterior margin; posterosuperior margin with single tooth ( Fig. 4H ). Metatibial surface finely microreticulate except extreme apex, and apex of lateral expansion shiny (probably from wear). Outer lateral metatibial margin serrate with row of small, setae-bearing teeth, posteromedian lateral projection large, shelf-like, apical margin uneven; row of more or less 3 tubercles anterior to lateral expansion; median surface and apex without tubercles; inner lateral margin with row of small, setabearing tubercles, emarginate in apical 1/3, small tooth present at base of emargination ( Fig. 4G ). Apex of 5th abdominal ventrite truncate. Genitalia long, dorso-ventrally flattened; basal piece shorter than parameres, proximal end curved; parameres straight in lateral view; median lobe longer than parameres, slightly wider at middle than a paramere, strongly curved upward in apical 1/3, apex rounded in ventral and lateral views ( Fig. 4F ).

Female. Apex of 5th abdominal ventrite same as in male.

Variation. Length 3.0 to 3.7 mm, width 1.7 to 2.0 mm. Carina on vertex may be absent; pronotal ridges vary somewhat in degree of spacing; surface sculpture varies slightly, clypeus and anterior portion of frons often shiny, probably because of wear; posterior trochanteral margin with 2 or 3 teeth, inner most tooth small, often not detectable.

Type material. Holotype male; Colorado: Hasty (Bent County), Colorado, 14.VI..82, Lot 11, BG& JL Carr ( JCIC) . Paratypes, 113: (46) same data as holotype except date 9.VI.83. Colorado: (1) Nunn, Pawnee Grassland , 6-VI-1977, J.W. Leetham; (24) New Mexico: Chaves Co., 35.5 mi. E. Roswell , VII-31- 1978; (2) Otero Co.; Three Rivers Petroglyphs , 7-VI-1987, Robert Gordon. Texas: (34) Hudspeth Co., 2mi N Fort Hancock, V-29-1982, V-30-1982, N. Rulien; (4) 10 mi. N. Andrews (Andrews Co.), 13.VI.83, Lot 2, BF& JL Carr; (1) Monahans (Ward Co.), 20.IV.87, BF& JL Carr. ( CNIC) ( JCIC) ( FSCA) ( MJPC) ( USNM) .

Other specimens. 18: (1) New Mexico: Eddy Co., 32 o 24.8’N, 103 o 41.5’W ; Eddy Co., 26 mi. N Carlsbad. (1) South Dakota: Angostura Dam, S. of Hot Springs, Vii 6-10, 1968, H.F. Howden. (1) Texas: El Paso Co. , 12 mi. NE Fabans, IV-23-1996, UV light, Coll. E. G. Riley-628; (2) Utah: (1) UTAH, Emery Co., Goblin Valley Rd .; Utah, Emery Co. 5100', 4airMiN GilsonBt, VII-20/23-81, Viers, Parker, Griswold ; Emery Co., Goblin Valley , 4-VIII-1986, Robert Gordon; (1) Utah, Emery Co., Little Gilson Butte , 29-VIII-1986, A. S. Menke; (4) Utah, Hanksville , VIII 7-9, 1968, H.F. Howden; (1) Utah, 14 mi. S. Hanksville , 25-VII-1968, J.E. Martin; (1) Fairview Ranch, 5000', 13 mi. S. Hanksville , 6.VIII.1968; (1) Utah, 26 mi. S. Hanksville (Garfield Co.), Hwy. 95, 16-VI-84, Robert Gordon ; Grand Co., 3 mi north of Moab, Hwy 123. ( BYUC) ( CMNC) ( CNIC) ( MJPC) ( TAMU) ( USNM) ( USUL) .

Remarks. In addition to key characters, G. costata differs from G. ecostata by the more roughly microsculptured body surface, distinct median pronotal furrow, bordered basal pronotal margin, and long metasternal setae.

Etymology. The name ecostata refers to a lack of elytral costae. Therefore, this species is named G. costata in reference to the slightly carinate appearance of the elytral striae, and in recognition of its close relationship to G. ecostata .

Figures

Figures 4A-4H. Glaresis costata Fall. A) Habitus view. B) Clypeus and head surface. C) Pronotal surface. D) Metasternum and mesotibia. E) Elytral striae. F) Male genitalia. G) Metaleg and trochanter, dorsal view. H) Metaleg and trochanter, ventral view.