Ambrysus pulchellus Montandon, 1909

Reynoso-Velasco, Daniel & Sites, Robert W., 2018, Revision of the Ambrysus pulchellus Montandon species complex (Heteroptera: Nepomorpha: Naucoridae), Zootaxa 4442 (3), pp. 351-377 : 361-372

Ambrysus pulchellus Montandon

( FIGs. 4–5 )

A. pulchellus Montandon 1897 : Verh. Zool.-Bot. Ges. Wien, 47: 11, 16–17 (key, original description).

A. pulchellus: Champion 1901 , Biol. Cent.-Amer., Insecta, 2: 356, Tab. 21 (list of taxa, illustrations).

A. pulchellus: Torre-Bueno 1906 , Proc. Entomol. Soc. Wash., 8: 54 (list of taxa).

A. nitidulus Montandon 1909 : Bull. Soc. Sci. Buc.-Roum., 17: 319, 326 (key, original description).

A. pulchellus: Montandon 1909 , Bull. Soc. Sci. Buc.-Roum., 17: 319 (key).

A. pulchellus: Kirkaldy & Torre-Bueno 1909 , Proc. Entomol. Soc. Wash., 10: 187 (list of taxa).

A. pulchellus pallidulus Montandon 1910 : Bull. Soc. Sci. Buc.-Roum., 18: 189–190 (original description) (NEW SYNONYM).

A. pulchellus: Van Duzee 1916 , Check List Hem., p. 52 (list of taxa).

A. p. pallidulus: Van Duzee 1916 , Check List Hem., p. 52 (list of taxa).

A. pulchellus: Van Duzee 1917 , Univ. Calif. Publ. Entom., 2: 458 (list of taxa).

A. p. pallidulus: Van Duzee 1917 , Univ. Calif. Publ. Entom., 2: 458 (list of taxa).

A. pulchellus: Hungerford 1920 , Univ. Kans. Sci. Bull., 11: 198, 200 (key, translation of original description).

A. p. pallidulus: Hungerford 1920 , Univ. Kans. Sci. Bull., 11: 198, 200–201 (key, translation of original description).

A. pulchellus: Millspaugh 1939 , Field and Lab., 7: 79 (list of taxa).

A. nitidulus: Usinger 1946 , Univ. Kans. Sci. Bull., 31: 187 (morphology).

A. pallidulus: Usinger 1946 , Univ. Kans. Sci. Bull., 31: 187 (species status).

A. pulchellus: Usinger 1946 , Univ. Kans. Sci. Bull., 31: 187 (morphology).

A. nitidulus: De Carlo 1950 , An. Soc. Cient. Argent., 150: 9 (list of taxa, distribution).

A. pulchellus: De Carlo 1950 , An. Soc. Cient. Argent., 150: 7 (distribution).

A. pulchellus: La Rivers 1951 , Univ. Calif. Publ. Entomol., 8: 303–306, 336 (supplemental description, habitus photo).

A. p. pallidulus: La Rivers 1951 , Univ. Calif. Publ. Entomol., 8: 306 (subspecies status, synonymy).

A. nitidulus: La Rivers 1953a , Univ. Kans. Sci. Bull., 35: 1305–1306 (synonymy).

A. pulchellus: La Rivers 1953a , Univ. Kans. Sci. Bull., 35: 1284, 1305–1308, 1342–1343 (key, excerpt from supplemental description, habitus photo).

A. pulchellus: La Rivers 1965 , Biol. Soc. Nev. Occas. Pap., 4: 6 (subgeneric assignment, distribution).

A. pulchellus: La Rivers 1971 , Biol. Soc. Nev. Mem., 2: 68 (list of taxa).

A. pulchellus nitidulus: La Rivers 1971 , Biol. Soc. Nev. Mem., 2: 68 (list of taxa, subspecies status) (NEW SYNONYM).

A. p. pallidulus: La Rivers 1971 , Biol. Soc. Nev. Mem., 2: 68 (list of taxa, subspecies status).

A. pulchellus: Davis 1986 , Southwest. Nat., 31: 538 (distribution).

A. pulchellus: Polhemus & Polhemus 1988 , Catal., p. 524 (list of taxa).

A. p. pallidulus: Polhemus & Polhemus 1988 , Catal., p. 524 (list of taxa).

A. p. pulchellus: Polhemus & Polhemus 1988 , Catal., p. 524 (list of taxa).

A. pulchellus: Sites & Bowles 1995 , J. Kans. Entomol. Soc., 68: 477 (distribution).

A. pulchellus: Davis 1996 , Southwest. Nat., 41: 3, 7–6, 13–14, 22–23 (key, diagnosis, biology).

A. p. nitidulus: Davis 1996 : Southwest. Nat., 41:13 (distribution).

A. p. pallidulus: Davis 1996 , Southwest. Nat., 41: 13, 22 (distribution, synonymy).

A. pulchellus : Herrera 2013, Dugesiana, 20: 226 (list of taxa, distribution).

A. p. nitidulus : Herrera 2013, Dugesiana, 20: 226 (distribution).

A. p. pallidulus : Herrera 2013, Dugesiana, 20: 226 (distribution).

A. pulchellus: Sites et al. 2018 , Aquatic Insects, 16–17 (distribution, key, diagnosis).

Diagnosis. ThIs spEcIEs cAN bE dIsTINGuIshEd fROm OThERs IN ThE cOmpLEx by ThE fEmALEs wITh ThE pOsTERIOR mARGIN Of ThE SGP NEARLy ROuNdEd; wITh ONLy A LOw cENTRAL cONcAvITy ANd INcONspIcuOus pOsTEROLATERAL cORNERs ( FIG. 4F ). ThE LATERAL mARGINs Of ThE SGP ARE NOT REfLExEd ANd ThE pOsTERIOR mARGIN Of LATEROsTERNITE VI Is sTRAIGhT ( FIG. 4F ). MALEs hAvE ThE AGP wIdEsT pREApIcALLy ( FIG. 4B ). ThE psEudOpARAmEREs hAvE A mEdIAL cONvExITy ANd TApER TO A NARROwLy ROuNdEd ApIcAL LObE ( FIG. 4C ). ThE scuTELLum hAs A NEARLy uNIfORmLy dARK bROwN cOLOR ( FIG. 4A ) ANd ThE dORsum Of ThE INsEcT Is NOTIcAbLy GLOssy.

Discussion. ThIs spEcIEs wAs dEscRIbEd bAsEd ON A sINGLE mALE spEcImEN cOLLEcTEd IN GuATEmALA; hOwEvER, NO spEcIfIc LOcALITy dATA wERE pROvIdEd. MONTANdON (1897) cONsIdEREd ThE sIzE ANd cOLOR Of ThE spEcIEs TO bE sImILAR TO ThOsE Of Ambrysus pudicus STåL. TwELvE yEARs LATER, MONTANdON (1909) dEscRIbEd A. nitidulus fROm GuAdALAjARA, MExIcO. MONTANdON cONsIdEREd A. nitidulus TO bE cLOsE TO A. pulchellus , buT sLIGhTLy LONGER ANd wITh wIdER EmbOLIA. AfTER ThE ExAmINATION Of spEcImENs fROm dIffERENT LOcALITIEs IN GuATEmALA, MExIcO, ANd ThE UNITEd STATEs, LA RIvERs (1953A) REcOGNIzEd A. pulchelus ANd A. nitidulus TO bE cONspEcIfIc ANd syNONymIzEd ThE spEcIEs, ARGuING ThAT ThE ObsERvEd vARIATION wAs TO bE ExpEcTEd bEcAusE Of ThE wIdE RANGE Of ThE TAxON. ALThOuGh LA RIvERs hAd syNONymIzEd A. nitidulus uNdER A. pulchellus , hE RETAINEd nitidulus As A subspEcIEs IN hIs cATALOG Of ThE fAmILy NAucORIdAE (LA RIvERs 1971), buT dId NOT pROvIdE AN ExpLANATION. WE hAvE ExAmINEd mORE ThAN 1,700 spEcImENs fROm EIGhT cOuNTRIEs ThROuGhOuT ThE RANGE Of ThIs spEcIEs; ALThOuGh vARIATION IN bOdy sIzE ANd EmbOLIum wIdTh Is EvIdENT, vARIATION IN mENsuRAL chARAcTERs Is NOT suffIcIENT TO suppORT ThE REcOGNITION Of subspEcIEs (sEE cOmmENTs IN ThE A. californicus DIscussION). FuRThER, ThE dIAGNOsTIc fEATuREs Of ThE spEcIEs ARE cONsTANT ThROuGhOuT ThE dIsTRIbuTION, wITh ONLy sLIGhT vARIATION. Thus, wE pROpOsE ThE subspEcIEs Ambrysus pulchellus nitidulus TO bE A juNIOR syNONym Of ThE NOmINATE subspEcIEs.

IN 1910, MONTANdON dEscRIbEd ThE subspEcIEs A. pulchellus pallidulus fROm TExAs, UNITEd STATEs. ThE dEscRIpTION wAs bAsEd ON spEcImENs cOLLEcTEd by GusTAv W. BELfRAGE ANd dEpOsITEd IN ThE USNM. MONTANdON sTATEd ThAT A. p. pallidulus dIffEREd fROm ThE NOmINATE spEcIEs IN ThE cOmpLETE AbsENcE Of dARK spOTs ON hEAd ANd pRONOTum. HE ALsO mENTIONEd ThAT ThE dARKER spEcImENs Of ThIs subspEcIEs wERE sImILAR TO hIs pREvIOusLy dEscRIbEd A. nitidulus . ALThOuGh NOT ExpLAINEd IN ThE ORIGINAL dEscRIpTION, IT Is LOGIcAL TO cONsIdER ThAT ThE spEcIfIc EpIThET pallidulus dERIvEs fROm ThE LATIN AdjEcTIvE " pallidus, " whIch mEANs "pALE," IN REfERENcE TO ThE pALE-yELLOw cOLORATION ON ThE hEAd ANd pRONOTum Of ThE subspEcIEs. AfTER ThE ExAmINATION Of spEcImENs cOLLEcTEd AT ROOsEvELT (TExAs), UsINGER (1946) NOTIcEd vARIATION IN cOLORATION, buT cONsIdEREd ThE wIdTh Of ThE EmbOLIA TO bE A cONsTANT fEATuRE ThAT cOuLd bE usEd TO dETERmINE ThIs TAxON; Thus, hE ELEvATEd ThE subspEcIEs A. p. pallidulus TO fuLL spEcIEs sTATus.

IN hIs wORK ON ThE Ambrysus Of ThE UNITEd STATEs, LA RIvERs (1951) dIsREGARdEd ThE TAxONOmIc chANGE mAdE by UsINGER (1946); ARGuING INTRAspEcIfIc vARIATION, hE pROpOsEd A. p. pallidullus As A syNONym Of A. pulchellus . LATER, LA RIvERs (1971) REINsTATEd ThE subspEcIfIc sTATus Of A. p. pallidullus wIThOuT fuRThER ExpLANATION. WE hAvE ExAmINEd ThE syNTypE spEcImENs usEd TO dEscRIbE ThE subspEcIEs, INcLudING ThE pALE cOLORATION ANd fAINT dARK mARKING ON ThE hEAd ANd pRONOTum, buT wE ATTRIbuTE ThEsE chARAcTERIsTIcs TO ThE cOLOR fAdING pROcEss. ThE dIffERENcE IN ThE EmbOLIA wIdTh Is subTLE ANd dOEs NOT REpREsENT AN INfORmATIvE chARAcTER upON whIch TO suppORT ITs sTATus As A subspEcIEs. ThE dIAGNOsTIc fEATuREs Of ThE spEcImENs ARE cONsIsTENT wITh ThOsE Of A. pulchellus ; bAsEd ON ThIs ANd fOLLOwING ThE sAmE LOGIc fOR ThE syNONymy pROpOsEd fOR A. p. nitidulus , wE pROpOsE Ambrysus pulchellus pallidulus ALsO TO bE A juNIOR syNONym Of Ambrysus pulchellus pulchellus .

Variation. SOmE fEmALEs pREsENT ThE pOsTERIOR mARGIN Of ThE SGP wITh mORE EvIdENT pOsTEROLATERAL cORNERs ANd cONspIcuOus cENTRAL cONcAvITy, Thus AppROAchING ThE fOuR-LObEd SGP Of A. californicus . SOmE mALEs cAN ExhIbIT AN AGP NOT wIdENEd pREApIcALLy, ThEREby dIspLAyING A NEARLy cONsTANT wIdTh IN ThE dIsTAL hALf.

Type locality. GuATEmALA.

Repository. HOLOTypE Is hOusEd AT ThE ZMH.

Distribution. Ambrysus pulchellus Is dIsTRIbuTEd fROm ThE sOuThERN UNITEd STATEs sOuTh TO NORThERN PANAmA ( FIG. 5 ). ThIs spEcIEs hAs ThE wIdEsT KNOwN dIsTRIbuTION AmONG spEcIEs IN ThE subGENus Ambrysus ANd pRObAbLy ThE ENTIRE GENus; ThE ONLy OThER spEcIEs wITh A sImILAR RANGE Is A. circumcinctus MONTANdON IN ThE subGENus Syncollus (sEE REyNOsO-VELAscO et al. 2016). AT ThE TImE DAvIs (1996) pREsENTEd hIs wORK ON ThE NAucORIdAE Of TExAs, hE pREsENTEd ThE dIsTRIbuTION Of A. pulchellus As RANGING fROm NORThERN GuATEmALA NORTh INTO cENTRAL MExIcO, ThEN NORThEAsTwARd INTO cENTRAL TExAs. IN TExAs, ThE NORThERNmOsT dIsTRIbuTION Of ThE spEcIEs wAs LOcATEd sOuThwEsT Of DALLAs; IN BOsquE, HILL, ANd SOmERvELL cOuNTIEs ( DAvIs 1996).

IN MExIcO ANd CENTRAL AmERIcA A. pulchellus Is dIsTRIbuTEd IN ThE ChIApAs ( GuATEmALA: BAjA VERApAz, GuATEmALA, IzAbAL, SuchITEpéquEz; HONduRAs: FcO. MORAzáN; MéxIcO: ChIApAs; NIcARAGuA: MATAGALpA) , BALsAs BAsIN (GuERRERO, MIchOAcáN, MORELOs, OAxAcA, PuEbLA) , MExIcAN GuLf ( BELIzE: CAyO DIsTRIcT; MéxIcO: OAxAcA, TAmAuLIpAs, VERAcRuz) , EAsTERN CENTRAL AmERIcA ( COsTA RIcA: CARTAGO; GuATEmALA: ZAcApA; PANAmá: ChIRIquí) , MExIcAN PAcIfIc COAsT ( COsTA RIcA: GuANAcAsTE; EL SALvAdOR: LA LIbERTAd, LA UNIÓN; GuATEmALA: SANTA ROsA; MéxIcO: ChIApAs, COLImA, GuERRERO, JALIscO, MIchOAcáN, NAyARIT, OAxAcA, SONORA; NIcARAGuA: LEÓN) , MExIcAN PLATEAu (COAhuILA) , SIERRA MAdRE dEL SuR (GuERRERO, OAxAcA) , SIERRA MAdRE ORIENTAL (QuERéTARO, SAN LuIs POTOsí, TAmAuLIpAs) , SONORAN (SONORA) , TAmAuLIpAs (NuEvO LEÓN, TAmAuLIpAs) , TRANsmExIcAN VOLcANIc BELT (JALIscO, MéxIcO, MIchOAcáN, PuEbLA) , ANd WEsTERN PANAmANIAN IsThmus ( COsTA RIcA: PuNTA ARENAs) bIOGEOGRAphIc pROvINcEs. IN MéxIcO ThE spEcIEs Is ALsO dIsTRIbuTEd IN ThE TRANsITION zONE bETwEEN ThE MExIcAN GuLf ANd SIERRA MAdRE ORIENTAL (HIdALGO, SAN LuIs POTOsí, TAmAuLIpAs, VERAcRuz) , bETwEEN ThE MExIcAN PAcIfIc COAsT ANd SIERRA MAdRE OccIdENTAL (NAyARIT, SINALOA) , bETwEEN ThE MExIcAN PLATEAu ANd TAmAuLIpAs (NuEvO LEÓN) , bETwEEN ThE SIERRA MAdRE OccIdENTAL ANd TRANsmExIcAN VOLcANIc BELT (JALIscO) , As wELL As ThE TRANsITION zONE bETwEEN ThE SIERRA MAdRE ORIENTAL ANd TAmAuLIpAs (NuEvO LEÓN) bIOGEOGRAphIc pROvINcEs. IN ThE UNITEd STATEs, ThE spEcIEs Is pREsENT IN ThE sOuThERN AREA Of ThE BAsIN ANd RANGE (ARIzONA) ANd ThE COAsTAL PLAIN (TExAs) , As wELL As ThE wEsTERN AREA Of ThE GREAT PLAINs (NEw MExIcO, TExAs) physIOGRAphIc pROvINcEs. IN MExIcO, IT Is wIdELy dIsTRIbuTEd sOuTh Of ThE MExIcAN PLATEAu bIOGEOGRAphIc pROvINcE ANd ExTENds NORThwARd INTO ThE sOuThERN UNITEd STATEs .

IN CENTRAL AmERIcA, ThE spEcIEs hAd pREvIOusLy bEEN REpORTEd fROm BELIzE, COsTA RIcA, EL SALvAdOR, ANd GuATEmALA. REpORTEd hERE ARE ThE fIRsT REcORds Of A. pulchellus fROm HONduRAs, NIcARAGuA, ANd PANAmA. HERRERA (2013) REpORTEd ThE spEcIEs fROm COsTA RIcA ANd EL SALvAdOR buT NO spEcIfIc dATA ON ThE LOcALITIEs wERE pROvIdEd. OuR REcORds REpREsENT ThE fIRsT spEcIfIc dATA fROm ThOsE cOuNTRIEs. Ambrysus pulchellus hAs bEEN REcORdEd IN ALL cOuNTRIEs IN CENTRAL AmERIcA.

BAsEd ON ThE dATA fROm ThE MATERIAL ExAmINEd ANd OuR ExTENsIvE cOLLEcTING, A. pulchellus Is ONE Of ThE fEw spEcIEs Of Ambrysus cOmmONLy fOuNd IN LENTIc hAbITATs. OThER sAucER buGs IN ThE NEw WORLd wITh pREfERENcE fOR sTILL OR sLOw-mOvING AquATIc hAbITATs ARE spEcIEs IN ThE GENus Pelocoris STåL.

Ambrysus pulchellus wAs cOLLEcTEd syNTOpIcALLy wITh A. abortus LA RIvERs, A. hungerfordi , ANd A. puncticollis STåL AT EL BAjíO IN JALIscO; wITh A. abortus , A. hungerfordi angularis LA RIvERs, A. melanopterus STåL, A. parviceps MONTANdON, ANd A. puncticollis AT ARROyO LOs ARRAyANEs IN JALIscO (L-1579); wITh A. ayoyolin REyNOsO & SITEs AT ITs TypE LOcALITy IN HIdALGO (L-1523); wITh A. buenoi IN Eddy COuNTy, NEw MExIcO (10-III-1989); wITh A. circumcinctus ANd A. lunatus IN SAN SAbA RIvER IN KImbLE CO., TExAs (21-V-1989); wITh A. circumcinctus , A. lunatus, Cryphocricos hungerfordi UsINGER, and Limnocoris lutzi LA RIvERs IN SOuTh LLANO RIvER IN KImbLE CO., TExAs (L-1645); wITh A. colimanus POLhEmus & POLhEmus AT ITs TypE LOcALITy IN COLImA (CL1229) ANd AT RíO ARmERíA (L-1571) IN COLImA; wITh A. contrerasi REyNOsO & SITEs AT ITs TypE LOcALITy IN MIchOAcáN (L-1486); wITh A. hungerfordi spicatus LA RIvERs AT PuENTE EL CARmEN (L-1318) ANd RíO LAGARTERO (L-1314) IN ChIApAs; wITh A. abortus , A. hydor LA RIvERs, ANd A. scalenus LA RIvERs AT ARROyO PAsO dE VIGAs IN ThE sTATE Of MExIcO (L-1909);

wITh A. lunatus AT OjO dE AGuA dE LAmpAzOs IN NuEvO LEÓN (19-XI-1985); wITh A. pudicus IN ThE sTATEs Of GuERRERO (24-VIII-1985) ANd MIchOAcáN (27-VIII-1985); wITh A. pudicus ANd A. signoreti STåL AT RíO CALAbAzO IN VERAcRuz (L-1524); wITh A. scalenus AT ITs TypE LOcALITy IN MIchOAcáN (28-VII-1936); wITh A. signoreti IN PuEbLA (L- 1417); wITh Cataractocoris shepardi SITEs, NOvELO & REyNOsO AT Is TypE LOcALITy IN ChIApAs (L-1359); ANd wITh Pelocoris b. biimpressus MONTANdON IN TwO LOcALITIEs IN TExAs: JAmEs RIvER IN MAsON CO. (24 OcT 1992) ANd A sTREAm NE Of WILLOw CITy IN GILLEspIE CO. (5 ApR 1997).

Published records. BELIzE: CAyO DIsTRIcT; COsTA RIcA; EL SALvAdOR; GuATEmALA: [EscuINTLA], [GuATEmALA], [IzAbAL], [BAjA VERApAz]; MéxIcO: ChIApAs, GuERRERO, JALIscO, [MéxIcO], MIchOAcáN, MORELOs, OAxAcA, TAmAuLIpAs; UNITEd STATEs: ARIzONA, NEw MExIcO, TExAs ( MONTANdON 1897, 1909, 1910; ChAmpION 1901; UsINGER 1946; LA RIvERs 1951, 1953A, 1971; DAvIs 1986, 1996; HERRERA 2013; SITEs et al. 2018). As A. nitidulus , ThE spEcIEs wAs REpORTEd fROm GuAdALAjARA ( MONTANdON 1909). As A. pulchellus pallidulus , IT wAs REpORTEd fROm TExAs ( MONTANdON 1910). HERRERA (2013) REpORTEd A. pulchellus fROm COsTA RIcA ANd EL SALvAdOR bAsEd ON dATA AvAILAbLE ON ThE INTERNET vIA ThE ELEcTRONIc CATALOG Of ThE ENTOmOLOGy COLLEcTION AT ThE CALIfORNIA AcAdEmy Of ScIENcEs; hOwEvER, wE dId NOT fINd ThE cORREspONdING spEcImENs duRING OuR vIsIT TO ThE CALIfORNIA AcAdEmy Of ScIENcEs.

Type material examined. HOLOTYPE ♂ (by mONOTypy): GUATEMALA: ALTE SAmmLuNG. / TypE / Ambrysus pulchellus MONTd. TypE / MONTANdON dETERm. SYNTYPES: MÉXICO: [Jalisco]: GuAdALAjARA, MExIcO, H ROLLE, BERLIN S.W. 11 / 6/24 / MusEum PARIs A.L. MONTANdON 1908 / A. nilidulus (sIc) MONTANdON TypE 1908 (1♀ MNHN); TypE / GuAdALAjARA, MExIcO, H ROLLE, BERLIN S.W. 11 / A. nitidulus MONLANdON (sIc) TypE 1908 / 6/24 / A.L. MONTANDON VEND. 25.11.1909 / BOx NR.: HET 52 (1Ƌ ZMH); GUADALAjARA, MExICO, H ROLLE, BERLIN S.W. 11 / 6/24 / A. nitidulus MONTANDON TYPE 1908 / J.R. DE LA TORRE BUENO COLLECTION K.U. (1♀ SEMC); GUADALAjARA, MExIcO, H ROLLE, BERLIN S.W. 11 / 6/24 / A. nitidulus MONT. / J.R. dE LA TORRE BuENO cOLLEcTION K.U. (1♀ SEMC). SYNTYPES: [UNITED STATES]: Texas: TExAs, BELfRAGE / FROm COLL U.S.N.M. / 0 58 / COTypE NO. 54269 U.S.N.M (1♀ USNM); SAME BUT / Ambr. pulchellus v. pallidulus MONTANDON, TYPE, 1909 (1♀ USNM); TExAS, BELfRAGE / FROm COLL U.S.N.M. / COTypE NO. 54269 U.S.N.M / Ambr. pulchellus v. pallidulus MONTANdON, TypE, 1909 / Ambrysus pulchellus MONTANDON 1897 (IRA LA RIVERS) (1Ƌ USNM); TExAS, BELFRAGE / COTYPE NO. 54269 U.S.N.M / Ambrysus pulchellus MONTANDON 1897 (IRA LA RIVERS) (2Ƌ USNM); TExAS, BELFRAGE / FROM COLL U.S.N.M. / COTypE NO. 54269 U.S.N.M (1♀ USNM).

Material examined. BELIZE: Cayo District: 3.1 Km S Of AuGusTINE, N16°57.041', W89°00.197', 456 m, 15 APRIL 2014 , COLL R.W. SITES & W.D. SHEPARD, SANDY/ROCKY STREAM wITH MARGINAL VEG, L-1726 (1♀ UMC); uNNAmEd sTREAm, 4.8 Km S Of AuGusTINE, N16°56.185', W89°00.581', 476 m, 15 ApRIL 2014, cOLL R.W. SITEs & W.D. ShEpARd, NARROw SANDY STREAM wITH OVERHANGING GRASSES, L-1727 (1Ƌ, 1♀ UMC); AUGUSTINE (5 KM N), 26 MAY 1986, COLLN 21, PAuL J. SpANGLER & RObIN A. FAITOuTE / cOLLEcTEd fROm RIO ON IN pOThOLEs, (2♂, 1♀ USNM). COSTA RICA: Cartago: TuRRIALbA, VII- 15–19-65, P.J. SpANGLER (1♀ USNM) . Guanacaste: GuANAcAsTE, 2 1957 JuL 12, D. LAucK / Ambrysus pudicus STåL 1862 ( IRA LA RIVERS) (1♀ UMC); 16 MI. S. LA CRUZ, VII- 13-1965 , PAUL J. SPANGLER (1♀ USNM). Punta Arenas: N. Of PALmAR NORTE, XII .28.1969, CL1279, J.T. POLhEmus / Ambrysus pulchellus MONT. dET. J.T. POLhEmus 1978 / C.J. DRAKE AccEssION (1♀ USNM). El SALVADOR: La Libertad: LAGuNA ChANmIcO, 6-X-76, H.C. ChApmAN / C.J . DRAKE AccEssION (1♀ USNM). La Unión: 15 MI. SW . LA UNION, VII- 31-1965, PAuL J. SpANGLER (1♀ USNM) . GUATEMALA: Baja Verapaz: LOs ENcuENTROs , 25 JuN 1966, FLINT & ORTIz (1♂ USNM). Guatemala: SAN JOsé PINuLA . 12 II 2001, R. dEL CId (1♀ UVG). Izabal: LOs AmATEs, KELLERmAN / MusEum Of COmpARATIvE ZOOLOGy / Ambrysus pulchellus MONTANdON dETERmINEd by IRA LA RIvERs 1948 (1♀ EMEC) ; LOs AmATEs, KELLERmAN / MusEum Of COmpARATIvE ZOOLOGy / Ambrysus pulchellus MONTANdON 1897 ( IRA LA RIvERs) (1♀ EMEC) . Santa Rosa: 1957 JUL 5, D. LAucK / Ambrysus pulchellus MONTANdON ( IRA LA RIvERs) / C.J . DRAKE AccEssION (1♀ USNM). Such[itepéquez]: EL ChOcLA [ChOcOLá], JuNE 11 1966, FLINT & ORTIz (1♀ USNM). Zacapa: GUALAN, GTA. [ PROBABLY 15-I-5] / J.R. DE LA TORRE BUENO COLLECTION K .U. (6Ƌ, 6♀ SEMC); SAME BUT 15-I-5 / C.J. DRAKE ACCESSION (2Ƌ, 1♀ USNM); SAME BUT / Ambrysus nitidulus MONT. K.U. '73 (1♂ USNM). Unknown: GuATEm. / MusEum Of COmpARATIvE ZOOLOGy / Ambrysus pulchellus MONTANdON 1897 ( IRA LA RIvERs) (1♀ EMEC) ; MINATITLAN [PROBABLY LAKE ATITLÁN] GTA / J.R. DE LA TORRE BUENO COLLECTION K.U. / C.J. DRAKE ACCESSION (1♀ USNM); SAME BUT / Ambrysus pulchellus nitidulus MONTd. (1♀ USNM). HONDURAS: Fco. Morazán: EL ZAmORANO, 850 m, N14°01', W87°00', MAyO–JuLIO [NO yEAR], EsTudIANTE EAP / 77.423 EAPZ (1♀ EAPZ); EL ZAMORANO, 850 M, N14°01', W87°00' , MAYO–JULIO [NO YEAR], ESTUDIANTE EAP / 77.425 EAPZ (1♀ EAPZ); 10 MI. N. SABANA GRANDE [SABANAGRANDE] , VII-29-1965, PAUL J. SPANGLER (2♀ USNM). MÉXICO: Chiapas: H[AcIEN]dA LA LIbERTAd , SEpT. 1 1937, H.D. ThOmAs / Ambrysus pulchellus MONTANdON 1897 (IRA LA

RIVERS) (1Ƌ EMEC); LA LIBERTAD, 1-1-38, 540 M, OCTAVIO UTRILLA LOUISE 500M ABOVE SEA LEVEL / 393 / Ambrysus pulchellus MONTANDON 1897 (IRA LA RIVERS) (1♀ SEMC); LA LIBERTAD, 1-1-38, 540 M, OCTAVIO UTRILLA LOUISE 500M AbOvE sEA LEvEL / 153 / Ambrysus pulchellus MONTANDON 1897 (IRA LA RIVERS) (1Ƌ SEMC); MPIO. ÁNGEL ALBINO CORzO, JALTENANGO, Km 3 cARR. JALTENANGO-REv. MExIcANA, 15°53'58.3''N, 92°43'48.3''W, 614m, L-1315, 7-JAN-12, SUBMERGED VEG., DRV COLL. (1Ƌ, 2♀ IEXA; 7Ƌ, 10♀ UMC); MPIO. ÁNGEL ALBINO CORZO, JALTENANGO, RÍO LAGARTERO, 15°52'33.7''N, 92°43'53'4''W, 605M, L-1314, 7-JAN-2012, SUBMERGED VEG., DRV COLL. (6Ƌ, 2♀ UMC); [MPIO. ChAmuLA], LAs OLLAs, 20 Km NE Of SAN CRIsTObAL dE LAs CAsAs ON ROAd TO TENEjApA, bOG, 6 MARch 1965, D.E. BREEDLOVE COLLECTOR (1Ƌ CAS); MPIO. CHIAPA DE CORZO, CARR. JALTENANGO-SALVADOR URBINA, 7 KM NW OF SALVADOR URBINA, 15°49'21.6''N, 92°47'15.2''W, 702 M, L-1344, 18-MAY-2012, DRV & PRH COLLS. (1♀ IEXA; 4Ƌ, 1♀ UMC); MpIO. ChILÓN, SANTA CRuz, RíO TuLIjA, 17°11'37.8''N, 91°44'11.6''W, 480m, L-1806, 01-JuN-2014, DRV & PRH COLLS. (1♀ IEXA); MPIO. HUIxTLA, HUIxTLA, RÍO HUIxTLA, 15°08'59.9''N, 92°27'04.6''W, 67 M, L-1901, 01-APR- 2015, DRV, SITES, SHEPARD, BARR & PRH COLLS. (1Ƌ, 1♀ IEXA); MPIO. JIqUIPILAS, RANCHO BONITO, PTE. RANCHO BONITO, 16°22'06''N, 93°52'04.2''W, 689M, L-1814, 03-JUN-2014, DRV & PRH COLLS. (1♀ IEXA; 2♀ UMC); MPIO. LA CONcORdIA, EL ALTILLO, RíO EL DORAdO, 16°06'27.5''N, 92°55'29.3''W, 524 m, L-1342, 18-MAy-2012, DRV & PRH COLLS. (1Ƌ IEXA); MPIO. MAPASTEPEC, MAPASTEPEC, RÍO NOVILLERO (RÍO SAN NICOLÁS), 15°25'20.9''N, 92°52'05.2''W, 37 M, L-1355, 21-MAY-2012, DRV & PRH COLLS. (2Ƌ, 6♀ UMC); MPIO. MARqUéS DE COMILLAS, EjIDO CHAjUL (BOCA dE ChAjuL), 16°06'47.9''N, 90°55'25.11''W, 171 m, L-1893, 30-MAR-2015, DRV, SITEs, ShEpARd, BARR & PRH cOLLs. (1♀ IEXA); MPIO. MARqUéS DE COMILLAS, SANTA RITA LA FRONTERA, PTE. MANZANERO, ARROYO MANZANARES, 16°04'59.1''N, 90°47'50.8''W, 182 M, L-1892, 30-MAR-2015, DRV, SITES, SHEPARD, BARR & PRH COLLS. (1Ƌ IEXA); MpIO. OcOsINGO, RANchO PuERTO RIcO, 16°04'46.9''N, 91°00'44.6''W, 183 m, L-1894, 30-MAR-2015, DRV, SITEs, SHEPARD, BARR & PRH COLLS. (1♀ IEXA); MPIO. SILTEPEC, MATASANO, RÍO MATASANO, 15°36'13.7''N, 92°33'37.1''W, 950 M, L-1348, 19-MAY-2012, DRV & PRH COLLS. (1♀ IEXA; 1Ƌ, 3♀ UMC); [MPIO. TEOPISCA], TEOPISCA, VII-21- 1964, PAUL J. SPANGLER (2Ƌ, 4♀ USNM); MPIO. TONALÁ, VELO DE NOVIA WATERFALL, 15°55'11.3''N, 93°29'45.9''W, 144 m, NR. PTE. LAs HERmANAs, L-1359 [TypE LOcALITy Of Cataractocoris shepardi ], 21-MAy-2012, DRV & PRH cOLLs. (1♀ UMC); MPIO. VILLA COMALTITLÁN, LÁZARO CÁRDENAS, KM 238 CARR. HUIxTLA-MAPASTEPEC, 15°10'47.4''N, 92°32'50.7''W, 16 M, L-1353, 20-MAY-2012, DRV & PRH COLLS. (1♀ UMC); MPIO. VILLA CORZO, JORGE ESPINOZA TORREs, ARROyO MAcOITE, 16°17'09''N, 92°59'32.6''W, 559m, L-1813, 03-JuN-2014, DRV & PRH cOLLs. (1♂, 2♀ IEXA; 3♀ UMC); MPIO. VILLA CORZO, PUENTE EL CARMEN, 16°07'51.4''N, 93°2'0.4''W, 547M, L-1318, 8-JAN-2012, SUBMERGED VEG., DRV COLL. (10Ƌ, 7♀ UMC); SAN CRISTOBAL, SEPT. 2 1937, H.D. THOMAS (1♀ SEMC); SAN VICENTE, 1-4- 38, 540 m, OcTAvIO UTRILLA L. / Ambrysus pulchellus MONTANDON 1897 (IRA LA RIVERS) (1♀ SEMC); SAME BUT / 154 (1Ƌ SEMC); 3 MILES SE OF COMOTAN [COMITÁN], 23 JUNE 1965, D.E. BREEDLOVE, CAL.ACAD.SCI.COLL. (1Ƌ, 2♀ CAS). Coahuila: [MpIO]. CuATRO CIéNEGAs, EL GARAbATAL [RIvER], COAh. MX., 9 JuLy 1999, COLL. A.E. COhEN, (1♂ TNHC); [MpIO. CuATRO CIéNEGAs], 3 mI N. CuATRO CIéNEGAs dE CARRANzA, C.J. McCOy, 30 MAR '64 / Ambrysus pulchellus MONT. DET. J.T. POLHEMUS / C.J. DRAKE ACCESSION (1Ƌ USNM); MPIO. VILLA DE JUARéZ, SECCIóN VEINTIDOS CERO, RÍO SALADO, 27°26'36.0''N, 100°28'38.9''W, 226M, L-1768, 18-MAY-2014, DRV & PRH COLLS. (8Ƌ, 5♀ IEXA; 20♂, 10♀ UMC). Colima: AmERIA [ARmERíA RIvER], CL1229 [TypE LOcALITy Of Ambrysus colimanus ], JTP, XI-26-68 / Ambrysus pulchellus MONT. DET. J.T. POLHEMUS '68 / C.J. DRAKE ACCESSION (1Ƌ USNM); MPIO. ARMERÍA, PERIquILLO, 3 Km E Of ARmERíA, RíO ARmERíA, 18°57'16.8''N, 103°56'45.7''W, 36 m, L-1571, 24-JuN-2013, DRV & PRH COLLS. (29Ƌ, 30♀ IEXA; 40Ƌ, 53♀ UMC); MPIO. COqUIMATLÁN, LOS AMIABLES, 19°09'48.9''N, 103°50'06.9''W, 288 M, L-1570, 23-JUN-2013, DRV & PRH COLLS. (2♀ IEXA; 1Ƌ, 1♀ UMC); MPIO. PISCILA, PTE. EL SALADO, RÍO SALADO, 19°11'39.8''N, 103°41'37.9''W, 371 M, L-1567, 23-JUN-2013, DRV & PRH COLLS. (6Ƌ, 8♀ IEXA; 4Ƌ, 4♀ UMC); 7 MI. E COLIMA, XII-22-1967, A. & A. GILLOGLY COLLECTORS (1Ƌ TAMU). Guerrero: Km 85 [cARR.] ZIHUATANEjO–LÁZARO CÁRDENAS, 27-VII-85, F. ARIAS (2♀ CNIN); SAME BUT / COLECCIóN DEL INSTITUTO DE BIOLOGÍA, UNAM. MéxICO D.F. (1♀ CNIN); KM 85 [CARR.] ZIHUATANEjO–LÁZARO CÁRDENAS, 27-VII-85, R. BARBA (1Ƌ CNIN); Km 95 CARRETERA COyucA dE CATALáN-ZIhuATANEjO, 24-VIII-85, F. ARIAs / COLEccIÓN dEL INsTITuTO dE BIOLOGíA, UNAM. MéxICO D.F. (1♀ CNIN); KM 145 CARR. IGUALA-CD. ALTAMIRANO, 23-XI-84, E. BARRERA / COLECCIóN DEL INSTITUTO DE BIOLOGÍA, UNAM. MéxICO D.F. (1♀ CNIN); KIL. 438 S OF MEx. CITY, H.D. THOMAS / Ambrysus pulchellus MONTANDON 1897 (IRA LA RIVERS) (1♀ SEMC); [MPIO. CHILAPA DE ÁLVAREZ], CHILAPA (GUERRERO), OCT. 29 1930, PROf. L. SchuITzE / Ambrysus pulchellus MONTANdON 1897 (IRA LA RIvERs) / IRA LA RIvERs cOLLEcTION BEqUEATHED TO THE CALIFORNIA ACADEMY OF SCIENCES - 1978 (1♀ CAS); MPIO. COYUCA DE CATALÁN, LAS CRUCES, RÍO LOS NOPALES, 18°14'49.6''N, 100°46'08.9''W, 314 M, L-1496, 28-DEC-2012, DRV & PRH COLLS. (1♀ IEXA; 2♀ UMC); [MPIO. COYUCA DE CATALÁN], PLACERES DEL ORO, 22-XI-84, H. BRAILOVSKY (1♀ CNIN); [MPIO. QUECHULTENANGO], COLOTLIPA, 17-I-86, COL. E. BARRERA (2♀ CNIN; 1♀ UMC); [MPIO. QUECHULTENANGO], JUxTLAHUACA, 26-VI-83, A. IbARRA / Ambrysus pulchellus MONTd. D.A. POLhEmus (1♂ CNIN). Hidalgo: MpIO. ATLApExcO, ATLALTIpA HuITzOTLAcO, PTE. ATLALTIpA, RíO VENAdO, 21°02'12.3''N, 98°22'24.7''W, 158m, L-1796, 27-MAy-2014, DRV & PRH COLLS. (1♀ IEXA; 1♀ UMC); [MPIO. MOLANGO], MOLANGO, LAGUNA DE ATEZCA (ARROYO), 13-MAR-1988, R. NOVELO COL. (2Ƌ IEXA); MPIO. MOLANGO, LAGUNA ATEZCA AT MOLANGO, VEGETATED MARGIN OF LAKE [ Hydrocotyle sp. pROb. ranunculoides L.f. ( ApIAcEAE )], 6 JuNE 2013, L-1519 / SITEs, DRV, PRH & ShEpARd, 20°48'27.9''N, 98°44'53.4''W, 1277M (5Ƌ, 5♀ IEXA; 21Ƌ, 12♀ UMC); MPIO. SAN FELIPE ORIZATLÁN, HUExTETITLA, ROCKS & ALGAE IN sLOw bEdROcK uNNAmEd sTRm., 7 juNE 2013, L-1525 / SITEs, DRV, PRH & ShEpARd, 21°09'44.4''N, 98°33'26.6''W, 154 M (2Ƌ, 1♀ UMC); MPIO. SAN FELIPE ORIZATLÁN, SAN FELIPE ORIZATLÁN, RÍO SAN PEDRO, 173 M, 21°10'13.6''N, 98°35'46.8''W, 7 JuNE 2013, L-1526 / SITEs, DRV, PRH & ShEpARd, bEdROcK wITh chANNELs, spARsE GRAvEL sTRIps, ALGAE, Elodea SP. (19Ƌ, 16♀ UMC); SAN FELIPE ORIZATLÁN, SAN FELIPE ORIZATLÁN, PTE. TUNTITLÁN, RÍO TUNTITLÁN, 21°10'03.4''N, 98°36'58.4''W, 175 M, L-1527, 07-JUN-2013, DRV, SITES, SHEPARD & PRH COLLS. (1Ƌ, 1♀ UMC); MpIO. TLANchINOL, cA. 2 Km W Of ChALchOcOTIpA, wATER sLIdE, 21°02'53.4''N, 98°35' 40.5''W, 681 m, L-1523 [TypE LOcALITy Of Ambrysus ayoyolin ], 06-JuN-2013, DRV, SITEs, ShEpARd & PRH cOLLs. (1♂, 1♀ UMC). Jalisco: MAGDALINA [MAGDALENA], 10-VII-1953, SLESNICK (1♀ USNM); MPIO. ATENGUILLO, EL BAjÍO, KM 55 CARR. MASCOTA- AyuTLA, cA. 2 Km NW LOs VOLcANEs, 20°20'08.2''N, 104°34'19.2''W, 1509 m, L-1581, 27-JuN-2013, DRV & PRH COLLS. (2Ƌ, 1♀ UMC); [MPIO. CIHUATLÁN], ZIHUATLAN [CIHUATLÁN], JALISCO, H. BRAILOVSKY, 12-I-75 (1♀ CNIN); MPIO. CIhuATLáN, CIhuATLáN, RíO MARAbAscO, 19°14'20.29''N, 104°33'11.22''W, 40 m, L-1572, 24-JuN-2013, DRV & PRH COLLS. (1♀ IEXA); [MPIO. GUADALAjARA], GUADALAjARA, 8-5-03 / J.R. DE LA TORRE BUENO COLLECTION K.U. (1♀ SEMC); MpIO. LA HuERTA, AGuA CALIENTE VIEjA dE ApAzuLcO, PTE. PuRIfIcAcIÓN, RíO PuRIfIcAcIÓN, 19°21'09.4''N, 104°53'22.8''W, 13 M, L-1573, 25-JUN-2013, DRV & PRH COLLS. (7Ƌ, 7♀ IEXA; 14Ƌ, 7♀ UMC); [MPIO. LA HuERTA], EsT. dE BIOLOGíA ChAmELA, 15-VII-1991, C. MAyORGA, G. ORTEGA / Ambrysus pulchellus MONTANdON dET. R. SITES 2009 (1♀ CNIN); MPIO. LA HUERTA, PTE. SAN NICOLÁS, RÍO SAN NICOLÁS, 19°39'20.8''N, 105°10'49.3''W, 11 M, L-1574, 25-JUN-2013, DRV & PRH COLLS. (1♀ IEXA); [MPIO. MAGDALENA], LA QUEMADA, DEC. 30 1958, MENKE & STANGE / Ambrysus pulchellus MONTANDON 1897 (IRA LA RIVERS) / C.J. DRAKE ACCESSION (1♀ USNM); [MPIO. MAGdALENA], LA QuEmAdA, DEc. 30 1958, MENKE & STANGE / Ambrysus pulchellus MONTANdON 1897 (IRA LA RIvERs) / IRA LA RIVERS COLLECTION BEqUEATHED TO THE CALIFORNIA ACADEMY OF SCIENCES - 1978 (2Ƌ, 1♀ CAS); MPIO. PUERTO VALLARTA, LAs PALmAs dE ARRIbA, 20°49'31.3''N, 105°05'25.3''W, 120 m, L-1578, 26-JuN-2013, DRV & PRH cOLLs. (1♀ UMC); MPIO. SAN SEBASTIAN DEL OESTE, PTE. LOS ARRAYANES, ARROYO LOS ARRAYANES, 20°49'44.5''N, 105°00'45.1''W, 337 M, L-1579, 26-JUN-2013, DRV & PRH COLLS. (3♀ UMC); MPIO. TOMATLÁN, TOMATLÁN, RÍO TOMATLÁN, 19°56'41.4''N, 105°14'36.1''W, 27 M, L-1575, 25-JUN-2013, DRV & PRH COLLS. (1Ƌ, 3♀ IEXA; 1Ƌ, 5♀ UMC); [MpIO. TOmATLáN], 2.3 Rd. Km E Of EL TuLE, sTREAm, 15 JAN. 2005, C.B. BARR / 20°00.98'N, 105°13.42'W, ELEVATION 210 FT. (1♀ EMEC); [MPIO. TOMATLÁN], 5.3 KM W EL TULE, 15 I 2005, 50', UNNAMED STREAM / WILLIAM D. SHEPARD, LEG (1♀ EMEC); [MPIO. TUxPAN], NR. ATENTIqUE [ATENqUIqUE], 3800', CL740, VI-12-1975, J.T. POLHEMUS / C.J. DRAKE AccEssION (1♀ USNM). México: MpIO. LuvIANOs, CAjA dE AGuA, 18°57'58''N, 100°17'30.9''W, 1079 m, L-1460, 20-DEC-2012, DRV & PRH COLLS. (1Ƌ, 1♀ IEXA; 3♀ UMC); MPIO. TEjUPILCO, PASO DE VIGAS, ARROYO PASO dE VIGAs, 18°53'32.6''N, 100°13'57.0''W, 1107 m, L-1909, 05-ApR-2015, DRV, SITEs, ShEpARd, BARR & PRH cOLLs. (4Ƌ, 4♀ IEXA; 7Ƌ, 6♀ UMC); PTE. LOS CUERVOS, 31-JAN-2008, K. CUEVAS & R. NOVELO COLS. (1Ƌ, 1♀ IEXA); TEjUPILCO, MEx. TEMESCALTEPEC [TEMASCALTEPEC], VI-18-33 / H.E. HINTON, R.L. USINGER COLLECTORS (1Ƌ EMEC); SAME BUT VI-19-33 (7Ƌ, 5♀ EMEC); SAME BUT / IRA LA RIVERS COLLECTION BEqUEATHED TO THE CALIFORNIA ACADEMY OF SCIENCES - 1978 (1Ƌ CAS); SAME BUT / Ambrysus pulchellus MONTANDON 1897 (IRA LA RIVERS) (1♀ CAS); TEjUPILCO, MEx. TEMESCALTEPEC [TEMASCALTEPEC], VI-22-33 / H.E. HINTON, R.L. USINGER COLLECTORS (1♀ EMEC); SAME BUT VI- 23-33 (1Ƌ EMEC); SAME BUT VI-30-33 (1Ƌ EMEC); SAME BUT / 301 / Ambrysus pulchellus MONTANdON 1897 (IRA LA RIvERs) (1♀ EMEC). Michoacán: EL SAbINO, URuApAN, MIch. MEx. [TypE LOcALITy Of Ambrysus scalenus ], 7-28- 36, H.D. ThOmAs / Ambrysus pulchellus MONTANDON 1897 (IRA LA RIVERS) (1♀ SEMC); SAME BUT / 392 (1♀ SEMC); EL SAbINO, URuApAN, MIch. MEx. 7-28-36, H.D. ThOmAs / Ambrysus pulchellus MONTANdON 1897 (IRA LA RIvERs) / IRA LA RIVERS COLLECTION BEqUEATHED TO THE CALIFORNIA ACADEMY OF SCIENCES - 1978 (1♀, 1? CAS); KM 95 CARRETERA ZIhuATANEjO-LázARO CARdENAs, 27-VIII-85, R. BARbA / COLEccIÓN dEL INsTITuTO dE BIOLOGíA, UNAM. MéxIcO D.F. (1♀ CNIN); LAGO [DE] PÁTZCUARO, III-90 (1Ƌ CNIN); [MPIO.] APATZINGÁN, CALIFORNIA, 07-JUN-2005, R. NOVELO COL. (1Ƌ IEXA); MPIO. AqUILA, EL DUÍN, PTE. OSTULA, RÍO OSTULA, 18°27'50.1''N, 103°32'34.1''W, 12 M, L-1489, 26-DEC- 2012, DRV & PRH COLLS. (2Ƌ, 3♀ IEXA; 4Ƌ, 4♀ UMC); MPIO. AqUILA, LA TICLA, PTE. IxTAPILLA, 18°24'56.6''N, 103°31'55.2''W, 0 M, L-1490, 26-DEC-2012, DRV & PRH COLLS. (1Ƌ, 1♀ IEXA; 2Ƌ UMC); MPIO. BUENAVISTA TOmATLáN, PIEdRAs BLANcAs, PTE. PIEdRAs BLANcAs, RíO GRANdE, 19°14'43''N, 102°46'16.9''W, 339 m, L-1482, 24-DEc- 2012, DRV & PRH COLLS. (2Ƌ, 2♀ IEXA; 6Ƌ, 1♀ UMC); MPIO. CARÁCUARO, CARÁCUARO, RÍO SAN DIEGO, 19°01'04.8''N, 101°07'33.2''W, 519 M, L-1473, 22-DEC-2012, DRV & PRH COLLS. (1Ƌ IEXA); [MPIO. COAHUAYANA], EL TICUIZ, 28-SEP-2005 (3Ƌ, 1♀ IEXA); [MPIO.] COAHUAYANA, EL TICUIZ (ARROYO), 10 MSNM, 20-APR-2006, R. NOVELO COL. (1Ƌ, 1♀ IEXA); MPIO. COALCOMÁN, EL SALITRE DE MARUATA, RÍO EL GAUYABAL, 18°52'50.4''N, 103°08'44.5''W, 1143 m, L-1486 [TypE LOcALITy Of A. contrerasi ], 25-DEC-2012, DRV & PRH COLLS. (1♀ IEXA); MpIO. JuáREz, cARR. ZITácuARO-TuzANTLA, N Of PARícuARO, 19°17'33.3''N, 100°25'53.4''W, 1327 m, L-1467, 21-DEc- 2012, DRV & PRH COLLS. (1Ƌ, 1♀ IEXA; 1♀ UMC); MPIO. LA HUACANA, LA HUACANA, COLONIA LÁZARO CÁRDENAS, 18°57'19''N, 101°48'43.2''W, 458 M, L-1478, 23-DEC-2012, DRV & PRH COLLS. (1Ƌ IEXA; 3Ƌ UMC); MPIO. MORELOs, LA PLAcITA dE MORELOs, PTE. LA PLAcITA, RíO AquILA, 18°32'19.9''N, 103°35'17.9''W, 7 m, L-1488, 26-DEc- 2012, DRV & PRH COLLS. (1♀ IEXA; 2♀ UMC); MPIO. TEPALCATEPEC, RÍO PINOLAPA (PUENTE), 28-MAR-2005, R. NOVELO COL. (1Ƌ IEXA); MPIO. TEPALCATEPEC, RÍO PINOLAPA, 10-JUL-2006, R. NOVELO COL. (1♀ IEXA); MPIO. TEpALcATEpEc, TEpALcATEpEc, 19°11'47.3''N, 102°50'44.9''W, 371 m, L-1483, 24-DEc-2012, DRV & PRH cOLLs. (2♂, 2♀ IEXA; 6Ƌ, 2♀ UMC); MPIO. TUZANTLA, TUZANTLA, RÍO TUZANTLA, 19°13'18.1''N, 100°34'05.5''W, 593 M, L-1470, 21-DEC-2012, DRV & PRH COLLS. (1Ƌ, 1♀ IEXA); PÁTZCUARO, [ILLEGIBLE MONTH]-31-38, H.D. THOMAS / Ambrysus pulchellus MONTANdON 1897 (IRA LA RIvERs) (1♀ SEMC). Morelos: [MpIO. CuAuTLA], CuAuTLA, 28-VIII-70, HARRy BRAILOvsKy cOL. / Ambrysus pulchellus MONTANDON 1897 (IRA LA RIVERS) (1♀ CNIN); [MPIO. CUERNAVACA], CUERNAVACA, APRIL 1959, N.L.H. KRAUSS, SEEPAGE POOL (1♀ CAS); MPIO. JOjUTLA, TEHUIxTLA, RÍO AMACUZAC, 18°33'41.8''N, 99°16'46.1''W, 856 M, L-1502, 31-DEC-2012, DRV & PRH COLLS. (2Ƌ, 1♀ IEXA; 2Ƌ, 1♀ UMC); [MPIO.] JOjUTLA, VICENTE ARANDA, RÍO AMACUZAC (900M), 11-DICIEMBRE-1982, P. LILjEHULT COL. (33Ƌ, 44♀ IEXA); [MpIO.] JOjuTLA, V[IcENTE] ARANdA, RíO AmAcuzAc, 27-MARzO-1983, V. GARcíA cOL., Ambrysus sp. R. NOvELO dET. 2004 (8Ƌ, 17♀ IEXA); SAME LOCALITY INFORMATION BUT 7-MAY-1983, G. RODRÍGUEZ COL. (9Ƌ, 5♀ IEXA); [MPIO.] TLAquILTENANGO, EL AsTILLERO, RíO CuAuTLA (850msNm), 05-FEbRERO-1983, I. OLIvA cOL. EN ORILLA c/vEGETAcIÓN (16♂, 9♀ IEXA); RIO AMACUZAC, MORELOS, KIL. 133 S. MEx. CITY, H.D. THOMAS / Ambrysus pulchellus MONTANdON 1897 (IRA LA RIVERS) (1♀ SEMC); S. RAFAEL A., 17-VI-82, M. ESPINOZA (11Ƌ, 13♀ CNIN); 4 MI N OF AMACUZAC, CL1039, 25 ApR 1964, J.T. & M.S. POLhEmus / Ambrysus pulchellus MONT. dET. J.T. POLhEmus / C.J. DRAKE ACCESSION (2Ƌ, 1♀ USNM); 14 MI. SOUTH CUERNAVACA, MEx. XII-8-48 / H.B. LEECH COLLECTOR / STOP #47 (1Ƌ, 1♀ CAS); 20 MI S OF CUERNAVACA, CL1047, 27 APR 1964, J.T. & M.S. POLHEMUS / C.J. DRAKE ACCESSION (1♀ USNM); sAmE buT / Ambrysus pulchellus MONT. dET. J.T. POLhEmus (1♀ USNM). Nayarit: [MpIO. COmpOsTELA], COmpOsTELA, DEc. 30 1958, MENKE & STANGE / Ambrysus pulchellus MONTANdON 1897 (IRA LA RIvERs) / IRA LA RIvERs cOLLEcTION BEqUEATHED TO THE CALIFORNIA ACADEMY OF SCIENCES - 1978 (2♀ CAS); MPIO. RUÍZ, VADO SAN PEDRO, RÍO SAN PEDRO, 21°57'01.7''N, 105°10'45.0''W, 13 M, L-1604, 02-JUL-2013, DRV & PRH COLLS. (1♀ UMC); [MPIO. SAN BLAS], ATICAMA, 5', CL727A, VI-7-1975, J.T. POLHEMUS / C.J. DRAKE ACCESSION (1Ƌ, 3♀ USNM); MPIO. STA. MARÍA DEL ORO, EL OCOTILLO, 21°13'46.4''N, 104°38'29.8''W, 1176 M, L-1605, 02-JUL-2013, DRV & PRH COLLS. (1Ƌ UMC); [MpIO. SANTA MARIA dEL ORO], TRIb. RíO GRANdE dE SANTIAGO, AbOuT 3 mI. N.E. Of Hwy 15 ON ROAd TO SANTA MARIA, MARCH 27 1955, M55-71 MILLER & GREENBANK (1♀ UMMZ); 26 MI. SE OF TEPIC, XI-23-1948 / H.B. LEECH (1Ƌ CAS). Nuevo León: MPIO. GARCÍA, RÍO PESqUERÍAS, 8-XI-85, S. TUFINIO (1♀ CNIN); SAME BUT / COLECCIóN DEL INSTITUTO DE BIOLOGÍA, UNAM. MéxICO D.F. (1♀ UMC); MPIO. GENERAL TERÁN, LA UNIóN, RÍO EL PILóN, 25°20'27.2''N, 99°34'57.5''W, 243M, L-1775, 19-MAY-2014, DRV & PRH COLLS. (3Ƌ, 2♀ IEXA; 6Ƌ, 3♀ UMC); MpIO. LAmpAzOs dE NARANjO, EL ANcÓN, RíO CANdELA, 27°02'33.5''N, 100°32'13.5''W, 293m, L-1770, 18-MAy-2014, DRV & PRH COLLS. (1♀ IEXA); [MPIO. LAMPAZOS DE NARANjO], LAMPAZOS, OjO DE AGUA, 19-XI-85 COL. R. BARBA, S. TUFINIO, H. ROjAS Y A. CONTRERAS / COLECCIóN DEL INSTITUTO DE BIOLOGÍA, UNAM. MéxICO D.F. (1Ƌ CNIN); MPIO. LOs RAmONEs, 1 Km N HAcIENdA EL CARRIzO, RíO SAN JuAN, 25°31'40.5''N, 99°37'20.9''W, 206m, L-1774, 19-MAy- 2014, DRV & PRH COLLS. (1Ƌ IEXA; 2Ƌ UMC); [MPIO.] SANTIAGO, LOS ATASCOSOS, 13-XI-85, R. BARBA (3Ƌ, 6♀ CNIN); SAME LOCALITY INFORMATION BUT S. TUFINIO (4Ƌ CNIN); [MPIO. SANTIAGO], LOS CAVAZOS, N.L., 15/V/74, A.M. GODÍNEZ (1Ƌ UMC); PUENTE CRISTOLINAS (SIC), 17 MI S OF MONTERREY, 19 JULY 1969, F.N. YOUNG #2303 / Ambrysus SP. ( HEMIPTERA : NAUCORIDAE ) (1Ƌ, 1♀ UMMZ); RÍO PILóN-N., VILLEGRAN (SIC), 19.JULY.1969, #2306 F.N. YOUNG (1Ƌ UMMZ); VILLA ALDAMA [VILLALDAMA], LA COYOTA, 13-XI-85, H. BARBA, (1Ƌ, 1♀ CNIN); SAME LOCALITY INFORMATION BUT H. ROjAS (2Ƌ, 1♀ CNIN); MPIO. VILLALDAMA, LA MORITA, RÍO SABINAS, 26°30'09.9''N, 100°18'24.5''W, 375M, L- 1772, 18-MAY-2014, DRV & PRH COLLS. (3Ƌ, 3♀ IEXA; 7Ƌ, 8♀ UMC); MPIO. VILLALDAMA, VILLALDAMA, RÍO SABINAS, 26°29'48.6''N, 100°25'50.4''W, 421M, L-1771, 18-MAY-2014, DRV & PRH COLLS. (1Ƌ, 1♀ IEXA; 1Ƌ, 2♀ UMC). Oaxaca: MpIO. MáRTIREs dE TAcubAyA, cARR. SAN FRANcIscO EL MAGuEy-SAN JuAN CAcAhuATEpEc, 10 Km SW Of SAN JuAN CAcAhuATEpEc, 16°33'57.1''N, 98°13'46.6''W, 207 m, L-1389, 29-MAy-2012, DRV & PRH cOLLs. (7♂ UMC); MpIO. MáRTIREs dE TAcubAyA, MáRTIREs dE TAcubAyA, 16°34'41.1''N, 98°13'08.7''W, 206 m, L-1390, 29-MAy- 2012, DRV & PRH COLLS. (8Ƌ, 3♀ IEXA; 5Ƌ, 3♀ UMC); MPIO. SAN ANTONIO NANAHUATIPAM, SAN ANTONIO NANAhuATIpAm, RíO SALAdO, 18°07'59.4''N, 97°07'42.6''W, 790 m, L-1877, 24-MAR-2015, DRV, SITEs, ShEpARd, BARR & PRH COLLS. (3Ƌ, 2♀ IEXA; 7Ƌ, 4♀ UMC); MPIO. SAN GABRIEL MIxTEPEC, KM 194 CARR. PUERTO ESCONDIDO- OAxAcA, SAN GAbRIEL MIxTEpEc, RíO RANA, 16°06'06.3''N, 97°03'52.9''W, 712 m, L-1380, 27-MAy-2012, DRV & PRH COLLS. (1Ƌ, 1♀ UMC); MPIO. SAN JOSé ESTANCIA GRANDE, 2 KM S OF SAN JOSé ESTANCIA GRANDE, 16°20'52.7''N, 98°14'07.5''W, 40 M, L-1386, 28-MAY-2012, DRV & PRH COLLS. (10Ƌ, 12♀ UMC); MPIO. SAN MIGUEL TLACAMAMA, SAN MIGuEL TLAcAmAmA, PTE. YuTANIñE, 16°25'32.6''N, 98°03'29.2''W, 260 m, L-1385, 28-MAy-2012, DRV & PRH COLLS. (9Ƌ, 10♀ UMC); MPIO. SAN PEDRO HUAMELULA, MORRO AYUTLA, PTE. RÍO AYUTLA, RÍO AYUTLA, 15°54'08.7''N, 95°51'13.3''W, 29M, L-1816, 4-JUN-2014, DRV & PRH COLLS. (1♀ IEXA); MPIO. SAN PEDRO MIxTEPEC, BAjOS DE CHILA, 15°54'44.4''N, 97°07'14''W, 20 M, L-1381, 27-MAY-2012, DRV & PRH COLLS. (40Ƌ, 30♀ IEXA; 32Ƌ, 44♀ UMC); [MpIO. SAN PEdRO TOTOLApA], TOTOLApAN [SAN PEdRO TOTOLApA]: RIO GRANdE, CL1063, 29 ApR 1964 J.T. & M.S. POLHEMUS / C.J. DRAKE ACCESSION (1Ƌ, 1♀ USNM); SAME BUT / Ambrysus pulchellus MONT. dET. J.T. POLhEmus (1Ƌ USNM); MPIO. SANTA MARIA ZACATEPEC, KM 189 CARR. STA. MARIA ZACATEPEC-PUTLA, S OF EL TAPANCO, 16°47'28.3''N, 97°59'20.4''W, 341 M, L-1393, 29-MAY-2012, DRV & PRH COLLS. (4Ƌ, 4♀ UMC); MPIO. SANTA MARIA ZAcATEpEc, Km 201 cARR. MáRTIREs dE TAcubAyA-SANTA MARIA ZAcATEpEc, cA. 13 Km NE Of SAN PEdRO AMUZGOS, 16°42'18.9''N, 98°00'52.9''W, 276 M, L-1391, 29-MAY-2012, DRV & PRH COLLS. (4Ƌ, 2♀ IEXA; 2Ƌ UMC); MpIO. SANTIAGO JOcOTEpEc, LA ALIcIA, 17°38'57.9''N, 96°00'38.7''W, 103 m, L-1369, 23-MAy-2012, DRV & PRH COLLS. (1Ƌ UMC); MpIO. SANTIAGO LLANO GRANdE, SANTIAGO LLANO GRANdE, 16°30'04.6''N, 98°17'55.2''W, 56 m, L-1388, 29-MAY-2012, DRV & PRH COLLS. (1Ƌ UMC); MPIO. VILLA DE TUTUTEPEC DE MELCHOR OCAMPO, KM 91 CARR. PuERTO EscONdIdO-PINOTEpA NAcIONAL, RíO GRANdE (PIEdRA PARAdA), RíO GRANdE, 16°00'38.6''N, 97°26'29.2''W, 22 m, L-1383, 28-MAY-2012, DRV & PRH COLLS. (1♀ UMC); TEHUANTEPEC, VII-15–16-1964, PAUL J. SPANGLER (3♀ USNM); 17 MI. E. JUCHITÁN, VII-5-1965, PAUL J. SPANGLER (13Ƌ, 8♀ USNM); 25 KM SE PINOTEPA NACIONAL, I-6-80 / REAREd sEEds NO. 1390-80 / REAREd sEEds Calopogonium mucunoides / C.D. JOhNsON cOLLEcTOR (1♂ TAMU). Puebla: MpIO. COxcATLáN, LA EsTAcIÓN, 5 Km SW Of COxcATLáN ANd 400 m W Of OLd TRAIN sTATION, 18°15'07.3''N, 97°11'24.5''W, 950 M, L-1876, 24-MAR-2015, DRV, SITES, SHEPARD, BARR & PRH COLLS. (1♀ IEXA; 1♀ UMC); [MPIO. IZÚCAR DE MATAMOROS], IZÚCAR DE MATAMOROS, VII-13-1964, PAUL J. SPANGLER (1♀ USNM); [MPIO. IZÚCAR DE MATAmOROs], MATAmOROs, 12 mI S.E. PuEbLA, MExIcO, VII-3-53, 3 700 fT / UNIv.KANs.MEx. ExpEdITION / SLEsNIcK NO. 31/ Ambrysus pulchellus MONTANDON 1897 (IRA LA RIVERS) (5Ƌ, 3♀ SEMC); SAME BUT / IRA LA RIVERS COLLECTION BEqUEATHED TO THE CALIFORNIA ACADEMY OF SCIENCES - 1978 (1♀ CAS); MPIO. JOLALPAN, JOLALPAN, RÍO NExAPA, 18°20'58.4''N, 98°49'18.2''W, 841 M, L-1501, 30-DEC-2012, DRV & PRH COLLS. (1♀ IEXA; 1♀ UMC); MPIO. TLAcuILOTEpEc, cARR. PLAN dE AyALA-TLAxcO, 2 Km E Of PApALOcTIpAN, 20°25'22.4''N, 97°56'38.2''W, 310 m, L-1417, 05-JUN-2012, DRV & PRH COLLS. (5♀ UMC); PUEBLA, 1957 JUN 27, D. LAUCK / Ambrysus pulchellus / Ambrysus pulchellus MONTANdON 1897 (IRA LA RIvERs) / C.J. DRAKE AccEssION (1♂ USNM). Querétaro: MpIO. ARROyO SEcO, Km 16 cARR. JALpAN-ARROyO SEcO, NR. EL TRApIchE, 21°19'56.3''N, 99°31'36.8''W, 650m, L-1757, 14-MAy-2014, DRV & PRH cOLLs. (1♂ IEXA). San Luis Potosí: MpIO. AquIsmÓN, LA CAñAdA (LOs CORRALEs), ARROyO EL NAcImIENTO, AppROx. 6 Km N Of TANchAchíN, 21°52'01.5''N, 99°09'04.5''W, 158m, L-1793, 24-MAy-2014, DRV & PRH cOLLs. (2Ƌ, 1♀ IEXA; 3Ƌ, 1♀ UMC); MPIO. AxTLA DE TERRAZAS, CóMOCA, RÍO TANCUILÍN, 21°25'02.9''N, 98°53'42.8''W, 86 M, L-1532, 08-JUN-2013, DRV, SITES, SHEPARD & PRH COLLS. (1Ƌ UMC); MPIO. CÁRDENAS, CANOAS, LA POZA AZUL, vEGTd. sTREAm bELOw dAm, 9 JuNE 2013, L-1536 / SITEs, DRV, PRH & ShEpARd, 21°57'04.4''N, 99°30'38.4''W, 969 m (4Ƌ UMC); [MPIO. CIUDAD VALLES], CIUDAD VALLES, CL1240, XII-15-69, J.T. POLHEMUS / C.J. DRAKE ACCESSION (1Ƌ USNM); sAmE buT / Ambrysus pulchellus MONT. DET. J.T. POLHEMUS '70 (1♀ USNM); MPIO. HUEHUETLÁN, EL NAcImIENTO dE HuIchIhuAyáN, 21°27'35.3''N, 98°58'38.8''W, 96 m, 8 JuNE 2013, L-1533 / SITEs, DRV, PRH & SHEPARD, SPRINGFED COLD STREAM wITH COBBLE (1Ƌ UMC); [MPIO. EL NARANjO], EL SALTO FALLS, 17 JUNE 1956, R.E. BEER & pARTy / Ambrysus pulchellus MONTANDON DET. J.R. DAVIS 94 (1♀ SEMC); MPIO. RAYóN, CANOAS, VEGETATED uNNAmEd sTREAm IN vILLAGE, 9 JuNE 2013, L-1537 / SITEs, DRV & ShEpARd, 21°56'30.9''N, 99°30'37.2''W, 999 m (4♂, 5♀ IEXA; 11Ƌ, 10♀ UMC); [MPIO. TAMAZUNCHALE], TAMAZUNCHALE, S.L.P., MExICO, VI-21-1953, 600 FT, SLESNICK NO. 11 / UNIv.KANs.MEx. ExpEdITION / Ambrysus pulchellus MONTANDON 1897 (IRA LA RIVERS) (1Ƌ, 1♀ SEMC); SAME BUT / IRA LA RIVERS COLLECTION BEqUEATHED TO THE CALIFORNIA ACADEMY OF SCIENCES - 1978 (1♀ CAS); MPIO. TANLAjAs, RíO COy AT MANANTIAL COy, 21°43'54.1''N, 98°58'35.4''W, 46 m, 8 JuNE 2013, L-1534 / SITEs, DRV & SHEPARD, VEGTD. STREAM MARGINS, ROOTMATS IN RIFFLE (3Ƌ, 3♀ IEXA; 5Ƌ, 5♀ UMC); [MPIO. TANLAjAS], RÍO COY (NACIMIENTO), 20-MAR-2006, R. NOVELO COL. (2♀ IEXA); PUENTE COVADONGA, KM 11 CARR. CD. VALLES-XILITLA (ARROYO 150MSNM), 04-JULIO-2004, R. NOVELO COL. (5Ƌ, 11♀ IEXA); 14 MILES EAST C[IUDAD] DEL MAIZ, S.L.P., MEx. XI-19- 48 / H. B. LEEch COLLEcTOR / IN sTREAm (1♀ CAS). Sinaloa: MpIO. ROsARíO, PTE. BALuARTE, RíO BALuARTE, 23°00'26.0''N, 105°51'04.5''W, 24 M, L-1599, 01-JUL-2013, DRV & PRH COLLS. (2Ƌ UMC); MpIO. SAN IGNAcIO, PTE. RíO PIAxTLA, RíO PIAxTLA, 23°53'23.8''N, 106°37'03.9''W, 46 m, L-1668, 22-DEc-2013, DRV & PRH cOLLs. (1♂ IEXA); 2 mI. E Of CONcORdIA, ARROyO EL ZApOTILLO, 20 JuLy 1952, JOhN D. LATTIN, CAL.AcAd.ScI.COLL. / IRA LA RIvERs cOLLEcTION bEquEAThEd TO ThE CALIfORNIA AcAdEmy Of ScIENcEs - 1978 (1♂ CAS). Sonora: MpIO. ÁLAmOs, EL NARANjO, ARROyO EL TAymucO, 27°14'43.3''N, 108°45'27.8''W, 462 m, L-1682, 25-DEc-2013, DRV & PRH cOLLs. (2♀ IEXA; 4♀ UMC); MPIO. ÁLAMOS, TAYMUCO (EL VADO), ARROYO LA VINATERÍA, 27°14'39.7''N, 108°43'17.3''W, 537 M, L-1683, 25-DEC-2013, DRV & PRH COLLS. (1Ƌ, 1♀ IEXA; 2Ƌ UMC); [MPIO. HERMOSILLO], HERMOSILLO, VII-9–16-53, B. MALKIN (12Ƌ, 19♀ CAS); [MPIO. HERMOSILLO], HERMOSILLO, VIII-15–20-53, B. MALKIN (2♀ CAS); [MPIO. HERMOSILLO], HERMOSILLO, V-26-55 / B. MALKIN COLLECTOR (1Ƌ, 1♀ EMEC); 20 MI. NNE CD. OBREGON, SON., VIII-20-69 / J.T. DOyEN COLLEcTOR / J. HAddOcK COLLEcTOR (1♀ EMEC). Tamaulipas: Km 63 [cARR.] Cd. VICTORIA–SOTO LA MARINA, 25-X-85, E. MEjORADA (1♀ CNIN); KM 91 CARR. TULA-CD. VICTORIA, 9-X-85, M. GARCIA / COLECCIóN DEL INSTITUTO DE BIOLOGÍA, UNAM. MéxICO D.F. (1Ƌ UMC); MPIO. ALDAMA, LAS NUBES, PTE. SAN RAFAEL, 23°09'56.4''N, 97°57'14.2''W, 37M, L-1789, 23-MAY-2014, DRV & PRH COLLS. (3Ƌ, 3♀ IEXA; 7Ƌ, 7♀ UMC); MpIO. JAumAvE, cA. 4 Km S Of JAumAvE (TOwARds PAdRÓN y JuAREz), RíO GuAyALEjO, 23°22'21.3''N, 99°23'47.5''W, 767M, L-1785, 22-MAY-2014, DRV & PRH COLLS. (2Ƌ, 2♀ IEXA; 2Ƌ, 2♀ UMC); MPIO. JIMéNEZ, LIBERTAD CAmpEsINA, PTE. LA LAjILLA, ARROyO LA LAjILLA, 24°08'48.5''N, 98°33'41.3''W, 114m, L-1778, 21-MAy-2014, DRV & PRH COLLS. (6Ƌ, 6♀ IEXA; 11Ƌ, 11♀ UMC); [MPIO. LLERA DE CANALES], LLERA TAMPS. MExICO, VI-15-53 / UNIv.KANs.MEx. ExpEdITION / Ambrysus pulchellus MONTANDON 1897 (IRA LA RIVERS) (4Ƌ, 4♀ SEMC); SAME BUT / IRA LA RIVERS COLLECTION BEqUEATHED TO THE CALIFORNIA ACADEMY OF SCIENCES - 1978 (1♀ CAS); MPIO. SAN FERNANDO, LAs NORIAs, ARROyO ChORRERAs, 24°36'30.0''N, 98°17'54.6''W, 70m, L-1777, 21-MAy-2014, DRV & PRH cOLLs. (2♂, 1♀ IEXA; 2Ƌ, 1♀ UMC); MPIO. SOTO LA MARINA, LAVADEROS, LAVADEROS DAM, 23°28'07''N, 98°01'05.4''W, 65M, L- 1787, 23-MAY-2014, DRV & PRH COLLS. (1Ƌ IEXA; 1Ƌ UMC); MPIO. SOTO LA MARINA, LOS PESCADOS, RÍO PALMAS, 23°37'20.3''N, 98°05'46.7''W, 24M, L-1786, 23-MAY-2014, DRV & PRH COLLS. (3Ƌ, 1♀ IEXA; 6Ƌ UMC); RUTA 101, 21-IV-85, COL. J. BUENO / COLECCIóN DEL INSTITUTO DE BIOLOGÍA, UNAM. MéxICO D.F. (1Ƌ CNIN); TAMPICO, MExIcO, JuLy 16 1950, DRAKE & HOTTEs / IRA LA RIvERs cOLLEcTION bEquEAThEd TO ThE CALIfORNIA AcAdEmy Of ScIENcEs - 1978 (2♀ CAS); 3 1–2 MI wEST FORLON TAMPS. AUG 8 1934, SMITH & DUNKLE / Ambrysus pulchellus MONTANdON DETERMINED BY IRA LA RIVERS 1948 (1Ƌ CAS); SAME BUT / H. CHANDLER COLLECTION (1♀ CAS); 3 1–2 MI wEST FORLON TAmps. AuG 8 1934, SmITh & DuNKLE / Ambrysus pulchellus MONTANDON (IRA LA RIVERS) (1Ƌ SEMC); SAME BUT / 150 (1Ƌ SEMC); 3 1–2 MI wEST FORLON TAMPS. AUG 8 1934, SMITH & DUNKLE / 151 / Ambrysus pulchellus MONTANDON (IRA LA RIVERS) (1Ƌ SEMC); 3 1–2 MI wEST FORLON TAMPS. AUG 8 1934, SMITH & DUNKLE / Ambrysus pulchellus MONTANDON (IRA LA RIVERS) (1♀ SEMC); 3 1–2 MI wEST FORLON TAMPS. AUG 8 1934, SMITH & DUNKLE / Ambrysus pulchellus MONTANdON dETERmINEd by IRA LA RIvERs 1948 / IRA LA RIvERs cOLLEcTION bEquEAThEd TO ThE CALIFORNIA ACADEMY OF SCIENCES - 1978 (1♀ CAS); 3 1–2 MI wEST FORLON TAMPS. AUG 8 1934, SMITH & DUNKLE / Ambrysus pulchellus MONTANdON 1897 (IRA LA RIvERs) / IRA LA RIvERs cOLLEcTION bEquEAThEd TO ThE CALIfORNIA ACADEMY OF SCIENCES - 1978 (2Ƌ, 1♀ CAS); 5 MI S CIUDAD VICTORIA TAMAULIPAS, MEx. 11-5-36, H.D. THOMAS / Ambrysus pulchellus MONTANdON 1897 (IRA LA RIvERs) / IRA LA RIvERs cOLLEcTION bEquEAThEd TO ThE CALIfORNIA AcAdEmy Of ScIENcEs - 1978 (1♀, 1? CAS). Veracruz: LA EspERANzA, RíO CAzONEs, 24-III-77, E. BARRERA / Ambrysus pulchellus MONT. DET. J.T. POLHEMUS (2♀ CNIN); MPIO. ATZALÁN, RÍO ALSESECA AT PUENTE TOMATA, 514 M, 19°55'15.7''N, 97°13'28.2''W, 13 JuNE 2013, L-1549 / SITEs, DRV, PRH & ShEpARd, ROcK, GRAvEL, ANd mARGINAL VEGTN. (1Ƌ, 2♀ UMC); [MPIO. CATEMACO], CATEMACO, J. BUENO, 9-V-81 (1Ƌ, 2♀ CNIN); SAME BUT / Ambrysus pulchellus DET. D.A. POLHEMUS (1♀ CNIN); [MPIO. CATEMACO], PRESA CFE CATEMACO, 8-V-81, ORTEGA (1Ƌ CNIN); [MpIO. CATEmAcO], PREsA CFE CATEmAcO, 9-V-81, M. GALIcIA / COLEccIÓN dEL INsTITuTO dE BIOLOGíA, UNAM. MéxICO D.F. (1Ƌ CNIN); [MPIO. CATEMACO], PRESA CFE CATEMACO, 9-V-81, R. NOVELO / COLECCIóN DEL INSTITUTO DE BIOLOGÍA, UNAM. MéxICO D.F. (1♀ CNIN); MPIO. IxHUATLÁN DE MADERO, NARANjO DULCE, RÍO VINAZCO, 20°49'24.9''N, 97°55'19.2''W, 86M, L-1798, 27-MAY-2014, DRV & PRH COLLS. (1♀ IEXA); MPIO. MISANTLA, CA. 2 Km E Of LIbERTAd, 20o03'59.5"N, 96o57'22.5"W, 56 m, L-1553, 14 JuNE 2013, DRV, SITEs, ShEpARd, NOvELO & PRH COLLS. (1♀ UMC); [MPIO. PAPANTLA], PAPANTLA, CL520, I-7-1971, M.S. & J.T. POLHEMUS / C.J. DRAKE ACCESSION (1Ƌ USNM); MpIO. PAsO dE OvEjAs, PAsO dE OvEjAs, RíO ATLIyAc, 19°17'04.1''N, 96°26'03.7''W, 40m, L-1802, 29-MAy- 2014, DRV & PRH COLLS. (1Ƌ IEXA); [MPIO. SAN ANDRéS TUxTLA], EST. DE BIOL. LOS TUxTLAS, 14-XI-85, E. RAMÍREZ

/ COLECCIóN DEL INSTITUTO DE BIOLOGÍA, UNAM. MéxICO D.F. (1Ƌ UMC); [MPIO. SAN ANDRéS TUxTLA, LOS TUxTLAS], SALTO DE EYIPANTLA, 6/V/81, G. ORTEGA LEóN (3Ƌ, 1♀ CNIN); MPIO. SAN RAFAEL, EL FAISÁN, 20°11'01.7''N, 97°05'38.5''W, 89 M, L-1547,12- JUN-2013, DRV, SITES, SHEPARD & PRH COLLS. (5♀ UMC); MPIO. STA MARIA, ROAD TO NARANjOs, 1 Km W Of IxcATEpEc, PTE. PEpEyucA, 11 JuNE 2013, L-1542 / 21°13'30.4''N, 97°59'28.8''W, 166 m, SITEs, DRV & PRH, ROCK TRICKLES BTw POOLS w/ VEG. & HEAVY ALGAE (4Ƌ, 3♀ IEXA; 9Ƌ, 8♀ UMC); MPIO. STA. MARIA, 4 Km W Of CITLALTEpEc, ROcKy sTREAm w/ ALGAE 11 JuNE 2013, L-1543 / SITEs, DRV, PRH & ShEpARd, 21°17'25.8''N, 97°54'56.8''W, 223 M (1Ƌ UMC); MPIO. TANTOYUCA, N OF PLATON SÁNCHEZ, EL REMANZO, RÍO CALABAZO, 21°17'08.2''N, 98°22'38.6''W, 41 M, L-1524, 06-JUN-2013, DRV, SITES, SHEPARD & PRH COLLS. (13Ƌ, 15♀ IEXA; 18Ƌ, 33♀ UMC); [MPIO. TANTOYUCA], 19 MI. SE TANTOYUCA, STREAM, I.8.1971, CL523 J.T. POLHEMUS / C.J. DRAKE ACCESSION (2Ƌ, 1♀ USNM); [MPIO.] TLAPACOYAN, EL ENCANTO, RÍO FILOBOBOS, 08-NOV-2009, R. NOVELO COL. (1♀ IEXA); [MPIO.] TLAPACOYAN, IxTACUACO (ARROYO), 30-MAR-2003, R. NOVELO COL. (3Ƌ, 2♀ IEXA); [MPIO.] TLApAcOyAN, IxTAcuAcO, 30-MAR-2003, Ambrysus SP., R. NOVELO COL. Y DET. (1Ƌ, 1♀ IEXA); S. OF GUTIERREZ ZAMORA / VERAcRuz, MExIcO, CL519, I-7-1971, J.T. POLhEmus / Ambrysus pulchellus MONT. dET. J.T. POLhEmus / C.J. DRAKE AccEssION (1♀ USNM). NICARAGUA: [León]: LEÓN, RíO POchOTE, 28-AbRIL-1993, R. NOvELO & J.M. MAEs cOLs. (1♂, 2♀ IEXA). Matagalpa: SELvA NEGRA, 12.998367°N, 85.91533°W, 1251m, 25–30 MAy 2012, K.B. MILLER ET AL; KBM 25051201 (1♀ UMC). PANAMÁ: Chiriquí: 3 Km NE CALdERA, 1 JuNE 1983, W.E. STEINER (2♂, 3♀ USNM). UNITED STATES: Arizona: COchIsE CO., ARIz., 7.29.1927, R.H. BEAmER / 322 / 90 / Ambrysus pulchellus MONT. (1Ƌ EMEC); PIMA CO. QUITOBAqUITO SPRING, ORGAN PIPE NAT. MON. / 1.IX.1961 COLLECTOR E. KIRSCHBAUM / C.J. DRAKE ACCESSION (1♀ USNM); SAME BUT Ambrysus pulchellus MONT. DET. J.T. POLHEMUS 1968 (1Ƌ USNM); PIMA CO., QuITObAquITO SpRINGs, ORGAN PIpE NAT'L MONu'T., AZ., 21-V-1971, R.L. SmITh / C.J. DRAKE AccEssION (1♂, 1♀ USNM); sAmE buT / Ambrysus pulchellus MONT. DET. J.T. POLHEMUS 1979 (1Ƌ USNM); [PIMA CO.], TUCSON, MAR. 21 1944, M.H. FROST JR. (1♀ USNM); [SANTA CRUZ CO.], PATAGONIA, CL 1217, III-25 -'67, J.T. POLHEMUS / Ambrysus pulchellus MONT. dET. J.T. POLhEmus '68 / C.J. DRAKE AccEssION (1♀ USNM). New Mexico: Eddy CO., 6.5 mI. S. 285 ON 137, 10 MARch 1989, cOLL: MORRIs & VAddER (1♂, 1♀ UMC). Texas: BLANcO CO., BLANcO RIv. AT BLANcO, 16 DECEMBER 1989, COLL: R.W. SITES (1♀ UMC); [BLANCO CO.], CYP[RESS] MILLS, TEx. / COLL. CHITTENDEN / Ambrysus nitidulus MONTD. DET. USINGER / C.J. DRAKE ACCESSION (1Ƌ USNM); BLANCO CO., PERDERNALES RIVER @ US281, 25 OCT 1992, S.K. JASPER (2♀ TAMU); BREwESTER CO., BIG BEND NAT'L PARK, BUTTRILL SPRINGS, 17 JULY 1991, R.C. VOGTSBERGER (1Ƌ UMC); COLORADO CO., CUMMINS CREEK, N29°45'56.7'' W96°33'3.4'', 20 JULY 1998, COLL. D.E. BOwLES (1♀ UMC); EDwARDS CO., NUECES RIV., 15 MI N. BARKSDALE, 7 DECEMBER 1989, COLL: J.A. BACK (1♀ UMC); GILLESPIE CO., LANGE'S MILL, APRIL 29 1971, V.V. BOARD / C.J. DRAKE ACCESSION (1Ƌ, 1♀ USNM); SAME BUT / Ambrysus pulchellus MONT. DET. J.T. POLHEMUS (1♀ USNM); GILLESPIE CO., PEDERNALES RIV. AT RT. 1, 4 MI E STONEwALL, 16 DECEMBER 1989, COLL: R.W. SITES (2Ƌ, 1♀ UMC); GILLESPIE CO., STREAM ON WILLOw CITY LOOP NE OF WILLOw CITY, 5 APR 1997, S.K. JASPER (1Ƌ, 1♀ TAMU); HAYS CO., SPRGS @ DEAD MAN'S HOLE, 12 NOV 1994, 30°19'N, 98°11'W, JASPER, GIBSON, & MOELLER (1♀ TAMU); [KERR CO.], NORTH FORK GUADALUPE RIVER, 0.5 MI. E OF HwY 39 ON 3140, 25 MAY 1997, COLLS: D WOODS & B NICHOLS (1♀ UMC); KERR CO., TEx., APRIL 9 1939, D. MILLspAuGh / Ambrysus pulchellus MONTANDON 1897 (IRA LA RIVERS) (1Ƌ EMEC); KIMBLE CO., BEAR CREEK, CA. 5 MI W JUNCTION, 10 FEBRUARY 1990, COLL: R.W. SITES (1♀ UMC); KIMBLE CO., BIG SALINE CK., 6.5 MI W LONDON ON WHISKEY RD., 8 JULY 1989, COLL: B.J. NICHOLS (1♀ UMC); SAME LOCALITY INFORMATION BUT 23 SEPTEMBER 1989, COLL: R.W. SITES (1Ƌ, 1♀ UMC); KIMBLE CO. JOHNSON FORK CK. AT SEGOVIA, 7 JULY 1988, COLL: R.W. SITES (1♀ UMC); SAME BUT 16 SEPT 1988 (1♀ UMC); KIMBLE CO., JOHNSON FORK CREEK, 15 DECEMBER 1989, COLL: R.W. SITES (1♀ UMC); KIMBLE CO., JUNCTION, LLANO R., 23-IX-1989, COLL: M. SUDDUTH (1♀ UMC); KIMBLE CO., JUNCTION-TTU CTR., SOUTH LLANO RIV., 14 DECEMBER 1988, COLL: R.W. SITES (1Ƌ, 1♀ UMC); KIMBLE CO., JUNCTION, SOUTH LLANO RIVER, 30°28'13.79''N, 99°47'04.93''W, L- 1645, 11 NOvEmbER 2013 / SITEs & REyNOsO-VELAscO, 515 m, ROcKy sTREAm wITh VEGETATED MARGIN IN SLOw CURRENT (1♀ UMC); KIMBLE CO., LLANO RIVER N. OF JUNCTION, APRIL 25 2001, G.L. CHALLET (1Ƌ UMC); KIMBLE CO., NORTH LLANO RIV., JUNCTION, 10 FEB. 1990, COLL: R.W. SITES (1♀ UMC); KIMBLE CO., Ox HOLLOw CREEK, CA. 8 MI W JUNCTION, 7 JULY 1989, COLL: R.W. SITES (6Ƌ, 9♀ UMC); KIMBLE CO., Ox HOLLOw CREEK, 7 MI WNW JUNCTION, 15 JULY 1989, COLLS: BAUGH, NICHOLS (1Ƌ UMC); KIMBLE CO., Ox HOLLOw CREEK, 9 MI WNW JUNCTION, 5 AUGUST 1988, COLL: W.R. SITES (8Ƌ, 8♀ UMC); [KIMBLE CO.], ROOSEVELT, TEx., IV-21-24 / PRES. BY J.O. MARTIN COLLEcTOR / 333 / Ambrysus pulchellus MONTANDON 1897 (IRA LA RIVERS) (1♀ EMEC); SAME BUT / Ambrysus pudicus STåL / Ambrysus nitidulus MONTD (1♀ EMEC); KIMBLE CO., SEGOVIA, JOHNSON FORK CK., 7 JULY 1988, COLL: R.W. SITES (1Ƌ UMC); KIMBLE CO., SOUTH LLANO RIVER AT TExAS TECH FIELD STATION, 30°28'22"N, 99°46'52"W, X- 14-2001, BAUMGARDNER, OSwALD & RILEY (1Ƌ TAMU); KIMBLE CO., SOUTH LLANO RIVER AT TTU CENTER-JUNCTION, 23

SEPT. 1989, COLLS: SITES & NICHOLS (3Ƌ, 2♀ UMC); SAME LOCALITY INFORMATION BUT 30 MARCH 1990, COLL: BACK AND BAUGH (1♀ UMC); [KINNEY CO.], SPRING FED STREAM W. OF BRACKETTVILLE, TEx., APR. 17 1939, H.H. & J.A. ROSS / INHS INSECT COLLECTION 651,930 (3♀ INHS); LEON CO ., HOTZEL RANCH, STREAM IN FLYNN BOG, 2 AUGUST 1991, COLL: S. JASPER (1Ƌ, 1♀ UMC); MASON CO., JAMES RIV., FM 2389, 10 MI. S MASON, 24 OCTOBER 1992, COLL: S. JASPER (1♀ UMC); MASON CO., JAMES RIVER NR. LONDON, 10 FEB. 1990, LOw wATER CROSSING, R.W. SITES (1Ƌ UMC); MASON CO., JAMES RIVER, 2 KM SW OF FM 2389, 24 OCT 1992, S.K. JASPER (5Ƌ, 7♀ TAMU); MCLENNAN CO., NORTH COw BAYOU @ OLD TEMPLE RD NEAR LORENA, 26 MAY 1993, S.K. JASPER (1♀ TAMU) ; MENARD CO., SAN SABA RIVER, 10 FEBRUARY 1990 , COLL: R.W. SITES (8Ƌ, 5♀ UMC); MENARD CO., SAN SABA RIVER AT MENARD, 4 AUG 1988 , COLL: R.W. SITES (1Ƌ, 2♀ UMC); SAME BUT 15 SEP 1988 (5Ƌ, 1♀ UMC); SAME BUT 15 DEC 1988 (8Ƌ, 4♀ UMC); SAME BUT 21 MAY 1989 (19Ƌ, 12♀ UMC); SAME BUT 9 JULY 1989 (11Ƌ, 5♀ UMC); SAME BUT 14 MAY 1990 (2Ƌ, 3♀ UMC); MENARD CO., SAN SABA RIVER @ MENARD, 21 MAY 1989 , COLL: T. DOEDERLEIN, IN RIVER EDGE (1Ƌ UMC); ROBERTSON CO., SOUTHwORTH BOG, 8 mI. E NEw BAdEN, JuLy 22 1970, V.V. BOARd / Ambrysus pulchellus MONT. J.T. POLhEmus / C.J. DRAKE ACCESSION (1♀ USNM); [TOM GREEN CO.], CHRISTOVAL, CONCHOS RIVER, CL1113, 9 MAY 1964, J.T. POLHEMUS / Ambrysus pulchellus MONT. DET. J.T. POLHEMUS / C.J. DRAKE ACCESSION (1Ƌ USNM); [VAL VERDE CO.], DEL RIO, TExAs, 7-8-38, R.H. BEAmER / Ambrysus pulchellus MONTANDON DETERMINED BY IRA LA RIVERS 1948 (1Ƌ EMEC) ; VAL VERdE CO., DOLAN FALLs, ROcK pOOLs w/ Chara @ DOLAN CK., 2 JUN 1997, S.K JASPER (1Ƌ, 1♀ TAMU); VAL VERDE CO., DOLAN FALLS PR., DOLAN CREEK, ALGAL MATS, 6 AUG 1994, S.K. JASPER (3Ƌ TAMU); VAL VERDE CO., DOLAN FALLS PR, SEEP NR DEVIL'S RIVER, 15 MAY 1996, S.K. JASPER (1♀ TAMU); VAL VERDE CO., DOLAN FALLS PR. VEG. @ SPRINGS @ DOLAN CK., 5 AUG 1994, S.K. JASPER (8Ƌ, 7♀ TAMU); VAL VERDE CO., DOLAN FALLS PR. VEG. @ SPRINGS @ DOLAN CK., 15 MAY 1996, S.K. JASPER (3Ƌ, 3♀ TAMU); VAL VERDE CO., DOLAN FALLS PRES., TNCT, PERM. POOL 1 MI N RIVER, 22 OCT 1994, S.K. JASPER (1♀ TAMU); VAL VERDE CO., DOLAN FALLS PRES., TNCT RD. xING @ Phragmites , 21 OcT 1994, S.K. JASPER (2♀ TAMU); VAL VERDE CO., DOLAN FALLS PRES., Tx. NATURE CONS., POOL IN DOLAN CREEK, OCT. 23 1994, S.K. JASPER (1♀ TAMU); VAL VERDE CO., DOLAN FALLS PRES., Tx. NATURE CONS. CANYON, POOLS AT DOLAN CREEK, VIII-6- 1994, S. JASPER (2Ƌ, 1♀ TAMU); VAL VERDE CO., 18 MI. S. JUNO DEVILS RIV. AT BAKER'S CROSSING, 6 AUG 1988, COLL: R.W. SITES (1Ƌ, 4♀ UMC); WILLIAMSON CO., EAST OF GEORGETOwN, SAN GABRIEL RIVER, 25 MARCH 1978, COLL: K. THOMSON (1♀ UMC); WILLIAMSON CO., SAN GABRIEL RIVER HwY 29, 4 APRIL 1987, COLL: T. BOTHwELL (1♀ UMC).

Figures

FIGURE 4. Structures of Ambrysus pulchellus. (A) dorsal habitus of female, (B) 6th and 7th abdominal terga of male, (C) 8th abdominal tergum of male, (D) ventral surface of phallosoma, (E) left side of phallososma, (F) sternum VI and subgenital plate of female (terminal abdominal segments removed). plc = posterolateral corner. Size bar = 0.5 mm and applies only to figures D and E.

FIGURE 5. Distribution of Ambrysus pulchellus in North America.