Colura section Gamolepis

Sangrattanaprasert, Jiroat, Chantanaorrapint, Sahut & Zhu, Rui-Liang, 2019, The genus Colura section Gamolepis (Lejeuneaceae, Marchantiophyta) in Malesian region, with the description of Colura sigmoidea, Phytotaxa 387 (1), pp. 40-54 : 41

Key to species of genus Colura section Gamolepis in Malesian region

1. Apex of lobule-sac acute, bearing apical crest (rarely without it)......................................................................................................2

- Apex of lobule-sac obtuse, lacking apical crest.................................................................................................................................4

2. Apical crest of the lobule sac consisting of 7–17 cells ........................................................................................................ C. cristata

- Apical crest of the lobule sac consisting of (1–)3–5 cells..................................................................................................................3

3. Lobule sacs ca. 1/2 as long as lobules; basal median cell of valve adnate with hinge cells forming a sigmoid curve in outline; dorsal margin of leaf lobe entire; female bracteole obtriangular, bilobed ................................................................................. C. sigmoidea

- Lobule sacs ca. 1/3 as long as lobules; basal median cell of valve adnate with hinge cells not forming a sigmoid curve in outline; dorsal margin of leaf lobe entire to crenulated; female bracteole subquadrate to semicircular, obtuse to emarginate .. C. verdoornii

4. Leaf lobes as wide as lobule sacs; underleaves 8–11 cells long and 4–5 cells wide ............................................................ C. meijeri

- Leaf lobes ca. 2 times width of lobule sac; underleaves 12–16 cells long and 6–9 cells wide............................................. C. valida