Drawida odaesan Blakemore, 2014

Blakemore, Robert J., Lee, Seunghan & Seo, Hong-Yul, 2014, Reports of Drawida (Oligochaeta: Moniligastridae) from far East Asia, Journal of Species Research 3 (2), pp. 127-166 : 152-153

23. Drawida odaesan Blakemore sp. nov.

Material. Holotype IV0000261295 (DNA sample HY1), mature specimen sketched and dissected, from Odaesan Mt. (alt. 1,563 m), Jinbu-myeon, Pyeongchang-gun, South Korea (ca. 37 47′53′′ N 128 32′35′′ E), collector Dr H-Y Seo 4 th July, 2013. Two immature Amynthas sp. from the same sample in a separate jar.

Etymology. After the type locality, noun in apposition.

Description. Slight blue-grey colour dorsally, distinctly demarcated cream ventrally. Prostomium proepilobous, peristomium ridged. Length 110 mm. Segments 210. Clitellum unclear. Dorsal pores absent. Nephropores well above d-lines towards mid-D on each side, most obvious from 12 onwards. Setae small and closely paired. Spermathecal pores in atria in 7/8 near c-lines. Male pores on 10/11 invaginated in bc-lines. Genital markings are postsetal, faintly on 7 above spermathecal pores; mid-ventral on 10 and paired on 11 in c-lines. Distinct equatorial indentations are on 10 and 11 either side of the male pores in 10/11. Female pores are paired anteriorly near b-lines in 12.

Septa 5/6-8/9 are thick. Low, sessile glands correspond to markings in 7 and much larger glands cover markings in 11 crossing 10/11 on each side (not present in mid-10). No atrium nor glands found near spermathecal pores in 8. Spermathecae on long, convoluted ducts in 8. Convoluted vasa deferentia go to body wall under GM glands on each side from testis sacs in 9/10, but no prostates present and penes if present, small. Ovisacs (empty) are elongated from ovaries in 11/12 extending to 16. Gizzards are in 13-17 (five of). Dorsal blood vessel is single but has distinctly arching vascular clusters in 18-21 and 24 and these are present in intervening segments and after 24 although not filled with blood. Nephridial bladders are elongate. Gut contains yellow soil without organic matter (geophagous).

Remarks. This species is close to D. nemora Kobayashi, 1938 that may also have five gizzards, but in 12-16 (as found in D. jirisanensis Hong, 2002 ) rather than 13-17 as here, and its GM markings at least also differ from both these other taxa. According to the DNA data (in Appendix, Fig. 1), Drawida odaesan has closest similarity amongst those tested to D. hattamimizu Hatai, 1930 which is interesting since it was considered a possible introduc- ed species to Japan. However, it not particular close genetically (just 80-81%) and it differs on many morphological points, such as size and GMs that, combined with the DNA barcode data (Appendix), serve to separate these taxa.