Drawida koreana nanjiro Blakemore, 2014

Blakemore, Robert J., Lee, Seunghan & Seo, Hong-Yul, 2014, Reports of Drawida (Oligochaeta: Moniligastridae) from far East Asia, Journal of Species Research 3 (2), pp. 127-166 : 145

14. Drawida koreana nanjiro Blakemore sub-sp. nov.

“ Drawida sp. 2 fat no penis NIBR ” in Blakemore & Lee

(2013: 128, fig. 1)

Material. Holotype (H) IV0000249928 (DNA sample WO 22) mature specimen, dissected, from NIBR facility Nanji-ro 42, Seo-gu, (GPS 37.564379, 126.638011), Incheon, collected by RJB 3 rd April, 2012 on path under pergola in rain, provisionally identified as “ Drawida cf koreana ” or “ Drawida sp. 2 fat no penis ”. Found with Amynthas carnosus (DNA WO 24) and the previous, Drawida koreana austri . Specimens IV0000261282 several matures possibly same from NIBR carpark collected in rain October, 2012 by RJB.

Etymology. Named after type locality (noun in apposition).

Description. Dark blue, especially the first 13 segments

Ovisac from 11/12

with darker dorsum and faint mid-dorsal line. Length 70 mm with 153 segments. Prolobous. Clitellum not developed. Dorsal pores absent. Nephropores in d-lines or above. Spermathecal pores in atria in 7/8 near c-lines and unilaterally in 6/7lhs (aberration). Male pores on 10 posteriorly in mid-bc-lines protrude on stubby mounds rather than penes. Female pores in b-lines on 12 near 11/12. Genital markings absent.

Spermathecal ducts moderately long to small atria with glands associated in 8. In 6/71hs is vestigial spermathecal aberration. Male pores on moderately long, coiled vasa deferentia have glandular prostates. Gizzards in 12-13, i.e., two of. Ovisacs present extending from 11 to segment 15. Gut has sandy quartz grits (geophagy).

Remarks. The current is similar to both Drawida koreana koreana Kobayashi, 1938 and D. japonica ( Michaelsen, 1892) , to the former in its blue colour and to the latter in its stubby penes and to both in its two gizzards in 12-13. It is especially close morphologically and is sympatric to the D. koreana austri Incheon paratype as well as to Drawida koreana shindo sp. nov. but is clearly separated from both by characteristics noted and marginally on its DNA data [in Appendix, Fig. 1 where BLAST WO22 (D. k. nanjiro) vs. WO27 (D. k. shindo) Identities=640/ 658 (97%), i.e., 3% different with 18 nucleotide bases exchanged; WO22 (D. k. nanjiro) vs. WO23 (D. k. austri) =596/660 (90%)]. Just 3% is a small genetic difference, here considered sufficient for sub-specific status due to concomitant morphological differences, as remarked on in the Methods section above. Full relationship to nominal D. koreana is pending confirmation of that taxon.