Drawida gisti gisti Michaelsen, 1931

Blakemore, Robert J., Lee, Seunghan & Seo, Hong-Yul, 2014, Reports of Drawida (Oligochaeta: Moniligastridae) from far East Asia, Journal of Species Research 3 (2), pp. 127-166 : 138

6. Drawida gisti gisti Michaelsen, 1931

Drawida gisti Michaelsen, 1931 a: 1 , 8, figs. 2-3; 1931b: 525. [From Tsinan, Shantung and Beijing, Hopei. Types in Hamburg (examined by Gates, 1940) and Beijing Museums].

Drawida gisti : Gates, 1935: 2; 1939: 406; 1940: 406; Kobayashi, 1937: 333, fig. 1; 1938: 85, fig. 2; 1940: 272; Blakemore & Kupriyanova, 2010: 16.

Distribution. Central and northern China and central and northern Korea (from Kobayashi, 1935: 95; 1940) abundant in Keijo (Seoul) and its vicinity and from Kinsen.

Description. Yellow grey, clitellum pinkish. Size 77-151 mm by up to 6 mm with 180-190 segments Prolobous. Setae small and closely paired (ab=cd). Dorsal pores absent. Clitellum 1 / 29,10 -13,14. Nephropores somewhat in line with c or more dorsal. Spermathecal pores 7/8 near to c-lines. Male pore in wide slits on blade-shaped penes in 10/11 b-c lines. Female pores paired on 12 near 11/ 12 in ab-lines. Genital papillae present, typically with a small pair, presetally on 10, or as larger discs in some of 7-12 with glands internally, or often absent.

Last hearts in 9. Spermathecal ampulla small with convoluted duct to ental end of large atrium in 7/8 with characteristic bulge, no accessory gland nearby. Testis sacs on 9/10 with coiled vas deferens entering ental end of large glandular prostate on penial pouch. Ovarian chamber in 10/11/12 with eggsacs extending to 13 or 14. Gizzards three in 12-14.

Remarks. Its subspecies D. gisti nanchangiana Chen, 1933: 200 is from China, the other sub-species, D. gisti anchingiana Chen, 1933: 202 was elevated to specific status by Kobayashi (1937: 333). Said by Gates (1940: 408) to be similar to D. hehoensis Stephenson, 1924 (from Myanmar) that was later associated with D. barwelli but differentiated on the greater size of the prostates and their relation to the penial pouches and the ‘urn-shaped’ spermathecal atrium plus GMs, when present. D. cheni Gates, 1935 was described by mistake by Chen (1933) as D. gisti f. typica which introduced some confusion. Genbank’s DNA barcode data in the Appendix has no close match.