Isoperla saccai (Festa, 1939)

Murányi, Dávid, 2011, Balkanian species of the genus Isoperla Banks, 1906 (Plecoptera: Perlodidae), Zootaxa 3049, pp. 1-46 : 4

saccai group

This group is characterized by triangular, elongate medial penial armature with an unfolded lateral edge that consists of long scales ( Consiglio 1967). The four species formerly included in the group: I. saccai ( Festa, 1939) , I. lugens , I. albanica Aubert, 1964 and I. russevi Sowa, 1970 are apparently not closely related and this group is probably polyphyletic. Isoperla albanica is transferred to a new group, the albanica group. Despite very different coloured armatures, the uncoloured structures of I. lugens are very similar to those of I. carbonaria (assigned to the acicularis group). SEM studies of additional species are needed to resolve the validity of these two groupings.