Cheilanthes

Kessler, Michael, Smith, Alan R. & Prado, Jefferson, 2017, Prodromus of a fern flora for Bolivia. XXVII. Pteridaceae, Phytotaxa 332 (3), pp. 201-250 : 207-208

Key to the Bolivian species of Cheilanthes

1 Segments abaxially densely scaly or hairy, the blade surfaces not visible ....................................................................................... 2

- Segments abaxially glabrous, glandular, farinose, or with sparse (to moderately dense in some forms of C. pilosa ) hairs or scales restricted to the veins ...................................................................................................................................................................... 10

2 Segments abaxially scaly .................................................................................................................................................................. 3

- Segments abaxially hairy .................................................................................................................................................................. 6

3 Segments adaxially glabrous (but partly hidden by curved scales from the abaxial surfaces); segments orbicular ........................ 4

- Segments adaxially sparsely hairy or with narrow hairlike scales; at least ultimate segments ovate to oblong .............................. 5

4 Rhizome scales concolorous, red-brown; blades 3-pinnate ............................................................................................ C. scariosa

- Rhizome scales bicolorous, dark brown with pale margins; blades 3-to 4-pinnate ................................... Myriopteris myriophylla

5 Rhizome scales more or less flat ................................................................................................................................... C. squamosa

- Rhizome scales with contorted apices ............................................................................................................................. C. incarum

6 Blades deltate, the lowermost pinnae the longest and with enlarged basiscopic segments ......................................... C. goyazensis

- Blades ovate to linear, the basal pinnae reduced .............................................................................................................................. 7

7 Rhizome scales pale orange-brown, concolorous; blades 2-pinnate or pinnate-pinnatisect ............................................ C. obducta

- Rhizome scales dark brown with pale margins or marginal setae; blades pinnate-pinnatifid to pinnate-pinnatisect ...................... 8

8 Hairs of abaxial surface of segments matted (tomentose), no or only a few hairs reaching beyond the segment margins; blades pinnate-pinnatisect ................................................................................................................................................ Myriopteris aurea

- Hairs of abaxial surface of segments long, evenly extending to or beyond segment margins; blades pinnate-pinnatifid ............... 9

9 Rhizome scales with abundant pale filamentous apices and marginal setae; basal segments usually the longest on a pinna; rachises lacking scales; veins on adaxial segment surfaces not visible ........................................................................................... C. fraseri

- Rhizome scales with few pale filamentous apices and marginal setae; basal segments usually slightly shorter than median segments on a pinna; rachises with a few deciduous scales among the abundant hairs; veins on adaxial segment surfaces impressed and visible ........................................................................................................................................................................... C. buchtienii

10 Rachises glabrous, glandular or farinose ........................................................................................................................................ 11

- Rachises, at least abaxially, with scales, gland-tipped hairs, or long, slender hairs ....................................................................... 15

11 Segments abaxially with sessile, red-brown glands ca. 0.1 mm long ..................................................................... C. rufopunctata

- Segments lacking glands ................................................................................................................................................................ 12

12 Indusia extending onto the segment stalks .............................................................................................................. Gaga marginata

- Indusia restricted to the segment margins ...................................................................................................................................... 13

13 Blades 1-pinnate-pinnatifid to 2-pinnate; ultimate segments lanceolate, acute; sori continuous along segment margins .................. ................................................................................................................................................................................... C. poeppigiana

- Blades 2-pinnate or more divided; ultimate segments ovate, obtuse; sori interrupted along segment margins ............................. 14

14 Rhizome scales concolorous, brown; blades 2-pinnate-pinnatifid to 4-pinnate; pinnules to 15 mm long ................. C. orbignyana

- Rhizome scales bicolorous, dark brown with pale margins; blades 2-pinnate; pinnules to 3 mm long ........ Adiantopsis tweediana

15 Blades linear, to 1.5 cm wide; ultimate segments linear, finger-like, to 1 mm wide ...................................................................... 16

- Blades deltate to lanceolate, 1.5–8 cm wide when expanded (may be narrower when dry and curled); ultimate segments ovate, 1.5–4 mm wide ............................................................................................................................................................................... 17

16 Blades pinnate to pinnate-pinnatifid; rhizome scales flaccid, orange-brown; pinnae with numerous, short, gland-tipped hairs ....... .................................................................................................................................................................................... C. micropteris

- Blades 2-pinnate-pinnatifid to 3-pinnate (barely 2-pinnate in undeveloped blades); rhizome scales rigid, dark red-brown; pinnae with a few hairs along the veins ...................................................................................................................................... C. pruinata

17 Rachises with both hairs and scales intermixed ............................................................................................................................. 18

- Rachises only with hairs ................................................................................................................................................................. 19

18 Rhizome scales castaneous; rachis hairs septate, pale red-brown with blackish cell boundaries; ultimate segments 1–3 mm long .. ........................................................................................................................................................................................ C. glutinosa

- Rhizome scales orangish; rachis hairs uniformly orangish; ultimate segments 2.5–6 mm long .......... Myriopteris notholaenoides

19 Rachises and blade hairs gland-tipped; rhizome scales dark brown ................................................................................... C. pilosa

- Rachises and blade hairs acuminate, lacking glands; rhizome scales orangish ........................................... Myriopteris moritziana