Harnischia tenuitubercula Chaudhuri & Chattopadhyay, 1990

Mukherjee, Bindarika & Hazra, Niladri, 2023, Taxonomic studies on Harnischia complex from India (Diptera: Chironomidae), Zootaxa 5278 (2), pp. 239-263 : 256-257

Harnischia tenuitubercula Chaudhuri & Chattopadhyay, 1990

Harnischia tenuitubercula Chaudhuri & Chattopadhyay, 1990: 158

Harnischia minuta Dutta & Chaudhuri, 1996: 272 , Syn. n.

Harnischia ohmuraensis Kobayashi & Suzuki, 1999: 80 , Syn. n.

Harnischia sibabecea Sasa, Sumita & Suzuki, 1999: 185 , Syn. n.

Material examined. Holotype male labelled ‘ H. sibabecea Sasa, Sumita & Suzuki, 1999 (http://www.type.kahaku. go.jp/TypeDB/diptera/973)’; Holotype male labelled ‘ H. minuta Dutta & Chaudhuri, 1996 (Type no. B. U. Ent. 210)’; 2 males, India, West Bengal, Purba Barddhaman, Burdwan [23.22°N, 87.85°E], 16.ii.2022, Coll. N. Hazra; 2 males, India, West Bengal, Purba Barddhaman, Burdwan [23.2446° N, 87.8496° E], 18.iv.2022, Coll. B. Mukherjee; 3 males, India, West Bengal, Jalpaiguri [26.71°N, 88.76 °E], 07.iv.2022, Coll. P. Hui’ .

Remarks. Harnischia ohmuraensis Kobayashi & Suzuki, 1999 , H. sibabecea Sasa, Sumita & Suzuki, 1999 , H. minuta Dutta & Chaudhuri, 1996 and H. tenuitubercula Chaudhuri & Chattopadhyay, 1990 have medially setose anal tergite, long gonostylus, broad anal point, and very small superior volsella. Kobayashi & Endo (2008) have synonymised H. sibabecea Sasa, Sumita & Suzuki, 1999 with H. ohmuraensis Kobayashi & Suzuki, 1999 . Besides that, Chen et al. (2017) mentioned that H. tenuitubercula Chaudhuri & Chattopadhyay, 1990 and H. ohmuraensis Kobayashi & Suzuki, 1999 may be conspecific. After a considerate study of types, we come to conclusion that H. minuta Dutta & Chaudhuri, 1996 is also conspecific with H. sibabecea Sasa, Sumita & Suzuki, 1999 . The original description of H. tenuitubercula was published in 1990 while descriptions of other species were made on 1996 and 1999 respectively. Therefore, the species H. minuta , H. ohmuraensis and H. sibabecea are junior synonyms of H. tenuitubercula .

A world key to the species of the genus Harnischia Kieffer (male)

(Modified after Chen et al., 2017)

1. Posterior margin of tergite IX concave at basal part of anal point (Palaearctic)...... H. quadricincta ( Goetghebuer, 1934b)

- Posterior margin of tergite IX not concave at basal region of anal point......................................... 2

2(1). Gonostylus short and broad............................................................................ 3

- Gonostylus long and roughly narrow................................................................... 14

3(2). Gonostylus with tooth-like or sharp projection at apex....................................................... 4

- Gonostylus without sharp projection..................................................................... 5

4(3). Gonocoxite with well-developed protuberance on distal inner margin; gonostylus with two bayonet-shaped projections (Palaearctic)..................................................................... H. cultriata Wang, 1999

- Gonocoxite without protuberance on inner margin; gonostylus with an inner apical tooth (Palaearctic and Orient)............................................................................. H. angularis Albu & Botnariuc, 1966

5(3). Gonostylus with inner basal expansion................................................................... 6

- Gonostylus without inner basal expansion................................................................ 8

6(5). Gonostylus with a strong bulbous-like inner expansion (Orient)..................... H. turgidula Wang & Jheng, 1993

- Gonostylus with weak inner expansion................................................................... 7

7(6). Anal point tapering, apex sharp (Orient, Palaearctic).................................. H. fuscimana Kieffer, 1921c

- Anal point parallel-sided, blunt and rounded apex (Palaearctic)....................... H. parallela Yan & Wang, 2016

8(5). Gonostylus expanded apically......................................................................... 9

- Gonostylus parallel-sided........................................................................... 10

9(8). Anal point tapered from base to apex; gonostylus with a little distal expansion (Palaearctic).................................................................................. H. biwacurtus Kawai, Okamoto & Imabayashi, 2002

- Anal point expanded in distal 1/3; gonostylus without a projection on distal part (Palaerctic).................................................................................................... H. japonica Hashimoto, 1984

10(8). Anal tergite band V shaped........................................................................... 11

- Anal tergite band Y shaped........................................................................... 12

11(10). Gonostylus highly curved in middle; anal point moderately long and broad (Orient, Holarctic, Australasia, Afrotropics)............................................................................ H. curtilamellata (Malloch, 1915)

- Gonostylus straight; anal point nearly short (Oriental China).................. H. longispuria Wang, Ji & Zheng, 1993

12(10). Anal point with pointed apex (Palaearctic)........................................... H. hachijoprima Sasa, 1994

- Anal point with blunt apex........................................................................... 13

13(12). Anal point tapered from base to apex (Palaearctic)........................... H. dissecta Zhang & Ferrington, 2017

- Anal point slender at base and wide in middle (Palaeartic)................... H. inaefeus Sasa, Kitami & Suzuki, 2001

14(2). Anal point very broad and long (Orient)........................ H. tenuitubercula Chaudhuri & Chattopadhyay, 1990

- Anal point moderately narrow in shape.................................................................. 15

15(14). Small lobe-like appearance lateral to anal point (Orient)........................................ H. bulbosa sp.n.

- Anal point without lateral lobe-like structure............................................................. 16

16(15). Anal point parallel in shape with blunt apex (Afrotropics)............................ H. lacteiforceps Kieffer, 1923

- Anal point constricted in middle with sharp pointed apex (Orient, Nearctic)................. H. incidata Townes, 1945