Eleutherogynium (Müll.Arg.) G.L.Webster ex R.W.Bouman 2018

Bouman, R. W., Keβler, P. J. A., Telford, I. R. H., Bruhl, J. J. & Welzen, P. C. van, 2018, Subgeneric delimitation of the plant genus Phyllanthus (Phyllanthaceae), Blumea 63 (2), pp. 167-198 : 170

Phyllanthus L. subgenus Gomphidium (Baill.) G.L.Webster section Adenoglochidion (Müll.Arg.) Müll.Arg. subsection Eleutherogynium (Müll.Arg.) G.L.Webster ex R.W.Bouman , stat. nov.

Phyllanthus L. subgenus Gomphidium (Baill.) G.L.Webster section Adenoglochidion (Müll.Arg.) Müll.Arg. subsection Eleutherogynium (Müll.Arg.) G.L.Webster ex R.W.Bouman. — Phyllanthus L. sect. Eleutherogynium Müll.Arg., Linnaea 32 (1863) 4, 14. — Type: Phyllanthus loranthoides Baill.

Glochidion J.R.Forst. & G.Forst. sect. Chorizogynium Müll.Arg. (1863) 58, 59. — Lectotype (designated by Webster 1986): Phyllanthus macrochorion Baill.

Phyllanthus L. sect. Heteroglochidion Müll. Arg. (1866) 319. — Type: Phyllanthus baladensis Baill .

Phyllanthus L. sect. Scleroglochidion Müll.Arg. (1866) 317. — Type: Phyllanthus myrianthus Müll.Arg .

Note — Section Scleroglochidion was previously placed in synonymy by Webster (1986) who expanded the description of Eleuterhogynium to include also Phyllanthus species with 3 free filaments. Section Heteroglochidion was defined by Müller on its biseriate sepals, which is a common character for subgenus Gomphidium . All of these sections are characterized by a rudimentary to absent nectar disc (see Müller 1866). Lobreau-Callen et al. (1988) in a palynological study, showed that the pollen of these groups showed a continuous variation in pollen characters and were difficult to differentiate. The lack of distinguishing floral and vegetative characters and the overlap in palynological characters leads us to the decision to combine the above sections in one subsection Eleutherogynium , with as main character the absent nectar disc to distinguish it from other species within section Adenoglochidion .