Physoglochidion (Müll.Arg.) R.W.Bouman 2018

Bouman, R. W., Keβler, P. J. A., Telford, I. R. H., Bruhl, J. J. & Welzen, P. C. van, 2018, Subgeneric delimitation of the plant genus Phyllanthus (Phyllanthaceae), Blumea 63 (2), pp. 167-198 : 170

Phyllanthus L. subgenus Gomphidium (Baill.) G.L.Webster section Gomphidium Baill. subsection Physoglochidion (Müll.Arg.) R.W.Bouman , stat. nov.

Phyllanthus L. subgenus Gomphidium (Baill.) G.L.Webster section Gomphidium Baill. subsection Physoglochidion (Müll.Arg.) R.W.Bouman. — Glochidion J.R.Forst. & G.Forst. sect. Physoglochidion Müll.Arg., Linnaea 32 (1863) 58. — Phyllanthus L. sect. Physoglochidion (Müll.Arg.) Müll.Arg., Prodr. 15,2 (1866) 318. — Type: Phyllanthus faguetii Baill.

Phyllanthus L. sect. Phyllocalyx Baill.(1862) 236 (nom.illeg., non Phyllocalyx A.Rich., 1847 ). — Glochidion J.R.Forst. & G.Forst. sect. Physoglochidion Müll.Arg. (1863) 58, 71. — Lectotype (designated here by R.W.Bouman, but see Webster (2001) manuscript synopsis of Gomphidium ): Phyllanthus faguetii Baill.

Note — Phyllanthus section Physoglochidion (Müll.Arg.) Müll.Arg. is characterized by 3 free stamens, 6 sepals in two whorls and a calyx that becomes saccate in fruit. Apart from the saccate calyx, these characters also occur in section Gomphidium and within section Physoglochidion and the saccate calyx shows a continuous variation between species ( Lobreau-Callen et al. 1988). Since these groups can also not be distinguished on palynological data we opt to reduce section Physoglochidion to a subsection level and place it with section Gomphidium .

Transfer of Phyllanthus oxycarpus to Glochidion :