Glenosema deki Azevedo & van Noort, 2019

Azevedo, Celso O. & Noort, Simon Van, 2019, Review of Afrotropical Glenosema Kieffer (Hymenoptera, Bethylidae) with description of 13 new species, Zootaxa 4585 (3), pp. 401-437 : 406-408

Glenosema deki Azevedo & van Noort , sp. nov.

Fig. 3

Description, holotype ♀. Body length 2.9 mm. Forewing length 0.1 mm.

Colour. Head black, mandible and antenna castaneous, palpi light castaneous; mesosoma black, except anterior part of pronotal collar and posterior margin of mesoscutellum dark castaneous, tegula castaneous, wings hyaline with vein light castaneous, legs castaneous, with tarsi light castaneous; metasoma with anterior terga almost black and posterior terga and sternites castaneous.

Head. Mandible 0.78 × LH, only slightly widened distally, cross-section thick and subtriangular, with seven distal teeth, lower tooth large, not visible in full frontal view, completely directed inward, three upper teeth in inclined series, dorsal margin placed at midheight of inner surface, very little denticulate on distal half, ventral margin not denticulate. Clypeus very short, its surface vertical, almost imperceptible in full dorsal view, triangular in frontal view, apical margin almost straight in frontal view, lateral lobe absent; median clypeal carina absent. Antennal foramina distant each other about 2.8 × antennal foramen diameter. First four antennomeres in ratio 22:12:5:5. Scape long and curved, evenly wide along its extension; pedicel longer than wide; flagellomeres about as long as wide, except last one distinctly longer than wide; flagellar pubescence dense, short and sub-erect. Eye with sparse and short setae, subtriangular, small, placed at dorsal half of head in lateral view. Frons coriaceous, punctures small, shallow, but conspicuous, irregularly distant one another; frontal line sulcate, short and inconspicuous. LH 0.82 × WH. WF 0.72 × WH. WF 3.92 × HE. OOL 1.9 × WOT. VOL 2.08 × HE. Distance of posterior ocellus to vertex crest 4.0 × DAO. Distance of anterior ocellus to supra-ocular line 3.0 × DAO. Frontal angle of ocellar triangle acute; ocelli tiny, hardly seen at median magnification, only slightly larger than punctures; ocellar triangle broad, POL 4.0 × DAO. Vertex almost straight medially. Sides of head posterior to eye parallel for short distance. Occipital carina complete. Malar space very small and triangular, eye almost touching dorsal mandibular condyle. Hypostomal carina evenly slightly curved, not angled medially.

Mesosoma. Coriaceous as frons. Dorsal pronotal area trapezoidal, wider than long; slope of pronotal collar almost vertical in profile. Mesonotum divided into anteromesoscutum and mesoscutellar-axillary complex, clearly separated by transscutal fissure. Anteromesoscutum very short, transverse, its surface flat in profile, about as long as mesoscutellum, without notauli and parapsidal signal. Mesoscutellar-axillary complex very short, transverse, its surface convex in profile, mesoscutellar sulcus almost crossing all anterior margin of mesoscutellum, conspicuous, deep, evenly slightly arched, not dilated laterally. Metanotum almost entirely covered by mesoscutellum. Metapectal-propodeal disc coriaceous, surface convex, conspicuously setose on anterior corner, median length 0.55 × median width, metapostnotal median carina conspicuous and complete; paraspiracular carina absent; lateral carina present and conspicuous; posterior carina complete; spiracle small, placed at dorsal surface attached to lateral carina. Propodeal declivity with its surface vertical and concave, median carina very inconspicuous, almost indistinguishable from surface. Prosternum small and concave. Mesopleuron with its surface strongly convex, dorsal area with subangularly produced protuberance. Micropterous, forewing reaching anterior transverse carina of metapectal-propodeal disc. Mesotibia subspinose. Tarsal claw arched with small medial tooth.

Metasoma. Slightly less coriaceous than frons. Metasomal tergite 3 with delicate median suture, carinate-like.

Variation. The body of the paratype is slightly shorter, with the head not as wide as in the holotype, but still much wider than long (LH 0.92 × WH).

Material examined. Holotype ♀, S. Africa, N. Cape, West Driefontein farm, Succulent Karoo 2 site (10.7 km 109° ESE Nieuwoudtville ), 31°22.581’S 19°13.507’E, Succulent Karoo on shales, 12–19 Oct 2000, HG Robertson, D Larsen, R Adams; Winkler, NW 00-SK2-W06 (SAM-HYM-P0024535) . Paratype: South Africa, Northern Cape: 1♀, Hantam National Botanical Garden, 741m, 31°24.182′S 19°08.587′E, 27 January–2 March 2009, S van Noort, Yellow pan trap, Nieuwoudtville Shale Renosterveld, GL 07-REN3-Y40 (SAM-HYM-P08745) ; 1♀, Tankwa National Park, Renoster River , 490m, 32°14.704'S 20°05.824'E, 9 Dec 2014 – 10 Jan 2015, S. van Noort, Malaise trap, Acacia karoo thicket, Tanqua Wash Riviere, Succulent Karoo, TKW14-ACA1-M14 (SAM-HYM- P08746) .

Diagnosis. This species differs from all other Afrotropical species of the genus by having a combination of several differences, the most evident are the wide head, the ocellar triangle not so far off the imaginary line between the dorsal eye margins, the mandibles in cross-section thick and subtriangular, and the metapectalpropodeal disc distinctly wider posterior with trapezoidal shape; and brachypterous.

Etymology. The specific epithet deki is an adjective from IsiXhosa, which means thick, in allusion to the mandible with thick cross-section.

Distribution. South Africa.

Figures

FIGURE 3. Glesonema deki sp. nov., holotype female. A. Habitus, lateral; B. Mesosoma, dorsal; C. Head, dorsal; D. Labels.