Gymnetis flavomarginata Blanchard, 1842

Iorio, Osvaldo Di, 2013, A review of the Cetoniinae (Coleoptera: Scarabaeidae) from Argentina and adjacent countries: systematics and geographic distributions, Zootaxa 3668 (1), pp. 1-87 : 29-31

Gymnetis flavomarginata Blanchard, 1842

Figs. 36–37 (aspects), 40 (parameres), 63 (distribution)

Gymnetis flavomarginata Blanchard, 1842 : plate 12, Fig. 1 ; Burmeister 1842: 800 (reference); Schaum 1844: 365 (observation).

Gymnetis flavomarginata Blanchard, 1847: 193 (description; distribution); Blanchard 1850: 34 (catalog; distribution); Burmeister 1861 (= 1943. Vol. I. 171: distribution); Schenkling 1921: 93 (catalog; distribution; references); Krajčík 1998: 77 (catalog; distribution); Antoine 2001: 122 (“ holotype ”, see Note 2 below), 124 (photo 1), 127 ( Figs. 8–9 ).

Gymnetis flavomarginata Blanchard , in part: Burmeister 1866: 576 (distribution).

Paragymnetis flavomarginata Blanch. subsp. flavomarginata Blanch. : Schürhoff 1937: 67 (new combination; distribution); Blackwelder 1944: 263 (catalog; distribution).

Distribution: BOLIVIA ( Burmeister 1866); ARGENTINA ( Schürhoff 1937, Blackwelder 1944, Krajčík 1998); Santiago del Estero ( Bruch 1911a); Corrientes: Caacaty [type locality], currently Nuestra Señora del Rosario de Caa Cati, January, D’Orbigny leg. [MNHN] (Blanchard [1847]); Corrientes, “Du voyage de M. d'Orbigny” [MNHN] ( Blanchard 1850); Corrientes ( Burmeister 1861); “Muséum Paris / Province de / Corrientes / d’Orbigny 1834”, “5731 / 94” [circular, handwritten], “732” [handwritten], “type” [red label, printed], “ G. / flavomarginata / Type Bl.” [handwritten], “ Gymnetis / flavomarginata / Blanch. / Holotype ” [red label, handwritten], 1 female [ MNHN] ( Antoine 2001); Santiago del Estero ( Bruch 1911a).

Material examined: BOLIVIA: Santa Cruz de la Sierra: “ S. Cruz / Boliv.” [printed on green paper], 1 male [ HBBA] ; PARAGUAY: Presidente Hayes: Costa Esmeralda, 23º 40’ S, 58º 29’ W (S23.66957, W58.49030), 18– 23-I-2011, U. Drechsel leg., 1 ex. [ ODIC]; ARGENTINA: Salta: Dto. Orán, Provincial Route 19, Km 4. (540 msm), 15-I-2002, J. Carreras leg., 1 male [ JCDT], 1 female [ ODIC ]; Santiago del Estero: “Rca. Argentina / Gob. Sgo. del Estero / 190_ / C. Bruch ”, 1 male [ CBBA], “ Gymnetis / pudibund a / Burm.” [handwritten by Bruch], “ Gymnetis / flavomarginata / Blanchard [1847] / Di Iorio det. 2012” ; Buenos Aires: “ Buenos / Aires ” [printed on green paper], Berg Collection, 1 ex. [ MLPA].

Male genitalia: parameres ( Fig. 40 ).

Note 1: Gymnetis flavomarginata was given as published in 1837 ( Krajčík 1998), 1837–1843 (Schenkling 1922), 1841 ( Burmeister 1842 [Fig.], 1866 [Fig.], Blanchard 1850, Antoine 2001), 1843 ( Bruch 1911 a, Blackwelder 1944), 1849 ( Morón et al. 2003), or without year ( Schürhoff 1937, Hardy 1975). According to Sherborn & Griffin (1934), plate 12 was published first (1842), and pages 185–232 were published later (1847). Therefore, the name was validated when it was first published in 1842 in association with an illustration. Accordingly, Burmeister (1842: 800) and Schaum (1844) cited only the figure of Blanchard, [1842], which was erroneously dated 1841 by Burmeister (1842).

Note 2: Blanchard (1847) gives the length of G. flavomarginata between 22 and 24 mm, an indication that he had more than one specimen (therefore a syntype series), but Antoine (2001) erroneously mentions that the type series consists of one female holotype (this specimen is a syntype).

Note 3: Burmeister (1866) identified three specimens, but only the specimen from Bolivia was correctly identified as G. flavomarginata . The two other specimens in his collection with the same identification proved to be G. bouvieri , one from Catamarca and the second from Santiago del Estero.

Note 4: Bruch (1911a) recorded G. flavomarginata from Santiago del Estero in his catalog, but the specimen in his collection was labeled as G. pudibunda . Santiago del Estero is not given for G. pudibunda in his list ( Bruch 1911a), nor is Corrientes for G. flavomarginata (Blanchard 1847) . In a letter to Bruch dated 13 October 1908, Moser mentioned two specimens of G. flavomarginata collected by Steinbach in Santiago del Estero that agree with the illustration of Blanchard (1842).

Note 5: The record of Buenos Aires may be due to mislabeling, but this is uncertain.

Note 6: The citations of G. flavomarginata from Mexico originated from the confusing nomenclature in the work of Schürhoff (1937), but should correspond to Gymnetis sallei Schaum, 1849 as was established by Antoine (2001). Morón et al. (2003) repeated the error.

Figures

FIGURE 1. Geographic distributions of Gymnetini from Argentina and adjacent countries: Blaesia atra (circles); Gymnetis bruchi (squares); Gymnetis cordobana (stars).

FIGURES 30–37. General aspects of Gymnetini from Argentina: 30–31, Gymnetis bouvieri (30, Catamarca; 31, Santiago del Estero); 32, Gymnetis carbo (Chaco); 33–35, Gymnetis cordobana (33, male; 34, female; 35, female, ventral view); 36–37, Gymnetis flavomarginata (36, male; 37, female).

FIGURES 8–12. Male parameres: 8–10, Gymnetis bajula (8, parameres; 9–10, details of the sculpture on the dorsal side in the angle between the parameres); 11–12, Gymnetis bruchi (11, parameres; 12, armature of the internal sac).

FIGURES 38–41. Male parameres: 38, Gymnetis bouvieri (Santiago del Estero); 39, Gymnetis cordobana (Córdoba: Villa Argüello); 40, Gymnetis flavomarginata (Bruch Collection); 41, Gymnetis carbo (Chaco: San Bernardo).