Elmomorphus elmoides, Selnekovič & Jäch & Kodada, 2024

Selnekovič, Dávid, Jäch, Manfred A. & Kodada, Ján, 2024, Taxonomic revision of the genus Elmomorphus Sharp, 1888. II. Redescription of the genus and review of the species from India, Nepal, Bhutan, Myanmar, China, Thailand, Laos, Cambodia, and Vietnam (Coleoptera: Dryopidae), European Journal of Taxonomy 957, pp. 1-229 : 51-55

Elmomorphus elmoides sp. nov.

urn:lsid:zoobank.org:act:483DEA09-BE29-47FC-8677-7F97D3224D41

Figs 21–22 , 28A , 109B

Differential diagnosis

Elmomorphus elmoides sp. nov. ( Fig. 21 ) can be characterised by the elongate and moderately convex body, widest behind the midlength of the elytra. The dorsal plastron is confined to the frontoclypeus, anterolateral portions of the vertex, and the lateral sides of the pronotum and the elytra. Elytra with small punctures scattered over the entire surface, and with large punctures arranged in longitudinal rows, which may sometimes be indistinct. Males with conspicuous long and erect setae or setal clusters on the labrum, prosternal process, and metaventrite. A similar combination of characters is shared with E. calvus sp. nov., which is larger, TL: ♂♂ 2.47–2.68 mm (2.55± 0.06, n=10), ♀♀ 2.65–2.94 mm (2.81 ± 0.09, n=10) in E. elmoides versus TL: ♂♂ 3.17–3.35 mm (3.25 ± 0.06, n=7), ♀♀ 3.15–3.45 mm (3.30 ± 0.11, n=10) in E. calvus sp. nov., and has a more convex body ( Fig. 23 ). In E. calvus , the pronotal plastron is confined to the anterolateral portions, while in E. elmoides it covers the entire lateral sides. The lateral pronotal sides are straight in E. elmoides but distinctly rounded in E. calvus . The phallobase is only slightly curved in E. elmoides ( Fig. 22 ) but strongly curved in E. calvus ( Fig. 24 ). The bursa copulatrix has large sclerotised spines ( Fig. 28A ), lacking in E. calvus ( Fig. 28B ).

Etymology

The epithet is a Latin adjective in the nominative singular referring to the superficial resemblance to various genera of Elmidae .

Type material

Holotype

VIETNAM – Lam Dong Province • ♂; “S-VIETNAM, 11°27′N 107°43′E, 15 km SW of Bao Loc , 900 m, 22.-24.IV.1995 P. Dembický ”; NMW.

Paratypes

VIETNAM – Lam Dong Province • 7 ♂♂, 6 ♀♀; same collection data as for holotype (on some labels “ Pacholatko & Dembicky ” are stated as collectors); CKB, NMW. – Gia Lai Province • 8 ♂♂, 6 ♀♀; “S-VIETNAM 40km NW An Khe Buon Luoi , 620-750m | 14°10′N 108°30′E 28.3. – 12.4.1995 Pacholatko & Dembicky ”; CKB, NMW .

Type locality

Vietnam, Lam Dong Province, 15 km southwest of Bao Loc, 900 m, 11°27′ N, 107°43′ E.

Description

Measurements (mm): TL: ♂♂ 2.47–2.68 (2.55 ± 0.06, n =10), ♀♀ 2.65–2.94 (2.81± 0.09, n=10); PL: ♂♂ 0.60–0.65 (0.63± 0.01, n =10), ♀♀ 0.65–0.73 (0.69± 0.03, n=10); PW: ♂♂ 0.98–1.06 (1.02 ± 0.03, n =10), ♀♀ 0.99–1.14 (1.08 ±0.04, n =10); EL: ♂♂ 1.85–2.05 (1.92± 0.06, n =10), ♀♀ 1.98–2.24 (2.12 ±0.09, n=10); EW: ♂♂ 1.17–1.32 (1.24± 0.04, n= 10), ♀♀ 1.24–1.43 (1.32± 0.06, n= 10); PhL: 0.92–1.01 (0.96 ± 0.03, n=10); 0.40–0.47 (0.43± 0.02, n= 10).

Body elongate-ovate ( Fig. 21 ), widest behind midlength of elytra, moderately convex dorsally, highest point at around midlength of elytra. Colouration black, except for reddish mouthparts, antennae, trochanters, and tarsi; remaining parts of legs brown to black. Dorsal vestiture consisting of short decumbent yellowish setae arising from small round punctures. Plastron on cranial surface except for posteromedian semicircular area on vertex, and along lateral sides of pronotum and elytra. Ventral surface covered with thin plastron setae, except on prosternal process, and median parts of metaventrite and ventrites 1–2.

Cranium dorsally covered with plastron, except semicircular area on vertex, and with round setiferous punctures, punctures slightly smaller than an eye facet, separated by 0.5–1.0 × puncture diameter. Labrum transverse, anterior margin shallowly emarginate medially; anterolateral angles rounded; surface microreticulate with small setiferous punctures; males with conspicuous long and erect setae, about as long as terminal maxillary palpomere, arranged in transverse row. Anterior margin of clypeus straight, with a row of short setae. Eyes large, oval, protruding, interfacetal setae short; ID: ♂♂ 0.30–0.42 mm (0.40 ± 0.01, n= 10), ♀♀ 0.41–0.47 mm (0.43± 0.02, n =10); APD/ID: ♂♂ 1.71–1.85 (1.80 ± 0.04, n =10), ♀♀ 1.67–1.80 (1.73 ±0.05, n= 10). Antennae short, 10-segmented, densely covered with plastron.

Pronotum transverse, widest at base, moderately convex, PW/PL: ♂♂ 1.56–1.76 (1.62± 0.06, n= 10), ♀♀ 1.50–1.64 (1.57± 0.04, n= 10). Surface smooth, with round setiferous punctures; plastron forming lateral band on each side, anteriorly about as wide as eye, constricted behind midlength.Anterior angles strongly deflexed, protruding and acute; pronotal sides convergent, straight. Hypomeron widest around midlength, covered with plastron. Prosternal process short and wide, lateral edges divergent and rounded; posterior edge rounded; lateral portions abruptly raised, wide, in males with cluster of long setae anteriorly; median portion flat. Scutellum longer than wide, smooth, with round setiferous punctures. Lateral parts of metaventrite covered with plastron; median part flat, smooth, with setiferous punctures, in males with two clusters of long setae posteriorly; metaventral process with lateral margins strongly raised, anterior margin weakly raised; discrimen apparent. Elytra ovate, widest behind middle, moderately convex, highest point around the middle; sides rounded, slightly diverging in anterior half and strongly converging in posterior half, EL/EW: ♂♂ 1.51–1.61 (1.54 ±0.03, n =10), ♀♀ 1.52–1.82 (1.61 ± 0.08, n =10); small punctures scattered, large punctures arranged in nine longitudinal rows (striae); strial punctures weakly delimited; plastron forming lateral band on each elytron extending from elytral side to fourth elytral stria. Tibiae straight; protibia ca 1.4 × as long as protarsus. Terminal protarsomere as long as three previous segments combined. Male foreclaws strongly curved, about half of length of terminal tarsomere, similar to female foreclaws.

Ventrites covered with plastron, except median portion of ventrites 1–2. Intercoxal process wide, subtriangular, with moderately raised margins and small round subapical depression; admedian keels moderately raised. Male ventrite 5 declivous near anterior margin; apex rounded, with several long erect setae medially. Female ventrite 5 evenly convex, with short longitudinal subapical keel, apex rounded. Aedeagus ( Fig. 22 ): phallobase long, moderately expanded proximally, PhL/PrL: 2.00–2.30 (2.21 ± 0.11, n= 10); parameres strongly curved ventrad in distal half, apices subacute (lateral aspect); penis apically relatively broad and rounded in lateral aspect, lateral extensions of apophyses strongly produced distad. (In most species of Elmomorphus , the apophyses are produced proximad and distad. In E. elmoides sp. nov. and E. calvus sp. nov., the apophyses are produced only distad. This character can be seen only in ventral view when the aedeagus is cleared in lactic acid and mounted on a slide.). Sclerotised fibula short and slender. Bursa copulatrix ( Fig. 28A ) with several large sclerotised spines; spermatheca large, tubular. Ovipositor heavily sclerotised; right coxite ca 1.4× as long as left coxite; valvifers ca twice as long as right coxite.

Secondary sexual dimorphism

Males possess clusters of long erect setae on labrum, prosternal process, and metaventrite. Ventrite 5 of female with longitudinal subapical keel.

Distribution

Vietnam ( Fig. 109B ).

Figures

Fig. 21. Elmomorphus elmoides sp. nov., paratype, male from Buon Luoi, Vietnam (CKB), TL: 2.55 mm.

Fig. 22. Elmomorphus elmoides sp. nov., paratype from type locality (CKB), aedeagus. A. Lateral aspect. B. Ventral aspect. Scale bar: 0.1 mm.

Fig. 28. Bursa copulatrix and vagina. A. Elmomorphus elmoides sp. nov., paratype from type locality (NMW). B. E. calvus sp. nov., paratype from type locality (CKB). C. E. depressus sp. nov., holotype (IAECAS). Scale bar: 0.1 mm.

Fig. 109. Distribution maps. A. Elmomorphus fusiformis sp. nov. B. E. elmoides sp. nov. C. E. calvus sp. nov. D. E. hongkong sp. nov. E. E. depressus sp. nov. F. E. sulcatus Kodada, Selnekovič & Jäch sp. nov.

Fig. 23. Elmomorphus calvus sp. nov., holotype, male (NMW), TL: 3.25 mm.

Fig. 24. Elmomorphus calvus sp. nov., holotype (NMW), aedeagus. A. Lateral aspect. B. Ventral aspect. Scale bar: 0.1 mm.