Laccophilus hendrichi, Hájek & Brancucci, 2015

Hájek, Jiří & Brancucci, Michel, 2015, A taxonomic review of the Oriental Laccophilus javanicus species group (Coleoptera: Dytiscidae), Raffles Bulletin of Zoology 63, pp. 309-326 : 314-319

Laccophilus hendrichi , new species

( Figs. 6 , 23 )

Type locality. Laos, Bolikhamsay province , Nam Kading NBCA [= National Biodiversity Conservation Area], Tad Paloy campsite.

Material examined. Holotype male, labelled: LAOS, Bolikhamsay Prov., / 18°23.17 N 104°09.65 E / NAM KADING NBCA , / Tad Paloy campsite , 300 m, / 8.-11. VII.2010, forest stream / Brancucci & Geiser leg. ( NHMB). Paratypes: 12 males, 10 females: same label data as holotype ( NHMB, NMPC); 77 males, 82 females: LAOS, Bolikhamsay prov., / Nam Kading NPA , Tad Paloy / campsite, 18°21-23'N, 104°09'E/,, 250-400 m, 24.-28. V. 2011 // NHMB Basel, / Laos 2011 Expedition, / M. Geiser, D. Hauck,/ A. Phantala & E. Vongphachan ( NHMB, NMPC); 1 male: LAOS: N-Vientiane Prov. / Vang-Vieng , 300 m / N 18°55′33′′, E 102°26′55′′ / 10-15.v. & 01-06.vi.2001 / JIŘÍ KOLIBÁČ leg. ( NHMB); 1 male: LAOS, Houa Phan Prov. / Ban Meuang Van Nera Muang Et , / 20°49-50‘N, 103°59‘-104°01-02‘E, / 2.-5.vi.2009, 300- 800 m, / Michael Geiser leg. // NHMB Basel, NMPC Prague / Laos 2009 Expedition: / M. Brancucci, M. Geiser, / Z. Kraus, D. Hauck, V. Kubáň ( NHMB); 66 males, 75 females: LAOS, Savannakhet prov., / Phou Xang He NBCA, ca. 5km / SW Ban Pa Phaknau, 250-400 m / 17°00‘ N, 105°38‘ E / 31.V.–6.VI. 2011 // NHMB Basel, / Laos 2011 Expedition, / M. Geiser, D. Hauck,/ A. Phantala & E. Vongphachan ( NHMB, NMPC); 12 males, 11 females: N Thailand-Road Mae Chem - / Mae Hong Son 4 km after Ban / Huai Boing , evergreen forest m / 1000 (m) 29.XII.1999 Mazzoldi ( NMPC, NHMW, PMCB) ; 2 females: Thailand: Mae Hong Son, Mae / Surin NP,, 2.ii.2012 ,, R. Müller / () // DNA / M. Balke / 5594 [respectively 5595] ( ZSMG) ; 1 male: Thai [ Thailand], 10- 16.5.1991 / CHIANG DAO [Chiang Mai Prov., Doi Chiang Dao, wildlife reserve], 600m / 19°24’N 98°55’E / David Král lgt. // Thailand’91 / “Thanon Thong Chai” / D. Král & V. Kubáň ( NMPC) ; 10 males, 8 females: Thailand bor. / Chiang Dao env. / 21.5.- 4.6.1995 / Snížek leg. ( JSCL, LHCM, NMPC) ; 1 female: THAILAND: 2.11.1995 / Chiang Mai Prov. / Doi Suthep, 750-800m / leg. Zettel (4) ( ZSMG) ; 1 female: THAILAND: 30.-31.11. / Chiang Mai Prov., 1995 / W Mae Rim, Mae Sa NP / leg. Zettel (2) ( NHMW) ; 2 males, 2 females: THAILAND: 16.11.1995 / Phrae Prov., Mae Khaem / 15km E Phrae , ca. 350 m / leg. Zettel (16b) ( NMPC, NHMW); 1 male: THAILAND: 24.11.1995 / Phetchanbun Prov. / Nam Nao NP / leg. Zettel (22) ( NHMW); 2 males: W – THAILAND 1990 / Tham Tharn Lot NP / N Kanchanaburi (3) / leg. Jäch 28.-29.11 ( NHMW) ; 1 male: W THAILAND: 300 m., / Thung Yai Wildlife / Sanctuary. / 15°28′N - 98°48′E. // Tak Province / Umphang District / Song Bae Stream. / 18-27.iv.1988. // Evergreen / rain forest. / M.J.D.Brendell. / B.M. 1988–183. ( BMNH); 2 males, 4 females: MYANMAR: Shan State / road Kalaw - Thazi , km 23 / 20°42.378′N 96°30.218′E / 720m, 24.10.1998 / leg. Schillhammer (30) ( NMPC, NHMW); 4 males: Myanmar (43) Shan State / NW Kalaw, km 23 on rd Kalaw- / Thazi , 20°42.378′N 96°30.218′E / rockpools, ca. 720 m, 16.5.1999 / leg. H. Schillhammer & R. Schuh ( NHMW) ; 3 males: Myanmar (136A) Shan State / Main Thouk Forest Monastery / 20°34.617′N 96°57.851′E / ca. 1160m, pools, 4.06.2004 / leg. Shaverdo & Schillhammer ( NHMW).

Additional non-type material. THAILAND: 1 male, Koh Samui , Hinta Hiu Yai, 8 January 1988, coll. Madl ( NHMW) ; 1 female, Sekhon Nakhon Province , Phu Pan NP, 490–520 m, 7–8 December 1995, coll. P. Schwendinger ( NHMW) . VIETNAM: 1 male, 12 km N Dalat, Lang Bian , 28–30 April 1994, coll. P. Pacholátko & L. Dembický ( NHMW) .

Diagnosis. Medium sized, oval oblong species (body length 3.8–4.4 mm). In habitus and shape of median lobe, the new species is similar to L. javanicus or L. tobaensis . However, L. javanicus has always reduced elytral colouration with anterior margin of the testaceous subbasal band straight, and apical part of median lobe in lateral view with slightly curved dorsal side. L. tobaensis is somewhat bigger (4.3–4.5 mm) and distinctly broader, with weakly impressed elytral sculpture.

Description. Colouration: Head and pronotum testaceous, elytra brownish-black with testaceous pattern consisting of sinuous subbasal transverse band well separated from base of elytra and interrupted on suture, indistinct mediolateral spot, sinuous preapical transverse band interrupted on suture, and a small indistinct apical spot ( Fig. 6 ) Appendages and ventral part testaceous.

Surface structures and sculpture: Head reticulation simple, composed of large irregular polygonal meshes, which are moderately deeply impressed. Frons between eyes, and clypeus with several large punctures. Reticulation of pronotum same as that of head. Pronotum along margins and in posterior angles with numerous large setigerous punctures. Reticulation of elytra simple composed of large irregular polygonal meshes similar to that of head or pronotum, but slightly more impressed. Meshes mostly complete, inner space often with several (1–3) small punctures. Each elytron with indistinct, sutural, discal and lateral row of large serial setigerous punctures. Ventral part reticulated with fine elongate meshes. Prosternal apophysis short and lanceolate, narrowly bordered. Abdominal ventrites with numerous oblique grooves and several large setigerous punctures, lateral margin bordered. Apical ventrite rounded posteriorly, with numerous large setigerous punctures and transverse grooves on disc. Male pro- and mesotarsomeres 1–3 weakly dilated with bundle of adhesive setae on ventral side. Claws of protarsi and mesotarsi simple and equal.

Male genitalia: Median lobe of aedeagus in lateral view as in Fig. 23 .

Female genitalia: Similar to male; pro- and mesotarsomeres not dilated.

Variability. Specimens of the type series vary considerably in extent and shape of the elytral colour pattern, especially those from Vietnam and Thailand (Koh Samui), which we therefore remove from the type series.

Measurements (N= 40): Body length 3.8–4.4 mm (mean value 4.2 mm, holotype 3.9 mm); width 2.2–2.5 mm (mean value 2.3 mm, holotype 2.2 mm).

Etymology. The new species is dedicated to our friend and colleague Lars Hendrich (Munich, Germany), a specialist on Dytiscidae .

Habitat. At the type locality, Laccophilus hendrichi was collected in small remnant pools of a drying up forest stream in rocky area (M. Geiser, pers. comm.).

Distribution. Laccophilus hendrichi occurs in (sub) mountainous areas of eastern Myanmar, northern Thailand and northern and central Laos ( Fig. 40 ).

Figures

Figs. 1–9. Habitus of Laccophilus: 1, L. aurofasciatus; 2, L. auropictus; 3, L. ceylonicus; 4, L. elegans; 5, L. girardi; 6, L. hendrichi, new species; 7, L. jaechi, new species; 8, L. javanicus; 9, L. komareki, new species. Scale bar = 1 mm.

Figs. 18–35. Median lobe in lateral view: 18, Laccophilus aurofasciatus; 19, L. auropictus; 20, L. ceylonicus; 21, L. elegans; 22, L. girardi; 23, L. hendrichi, new species; 24, L. jaechi, new species; 25, L. javanicus; 26, L. kalimantanensis, new species; 27, L. komareki, new species. 28, L. mahakamensis; 29, L. mazzoldii, new species; 30, L. nusatenggaraensis, new species; 31, L. schillhammeri, new species; 32, L. stastnyi, new species; 33, L. tobaensis; 34, L. tonkinensis; 35, L. wittmeri. Scale bar = 0.5 mm.

Figs. 40–41. Distribution of the species of the Laccophilus javanicus group