Mallochia aurantia ( Santos & Aguiar, 2008 ) Supeleto & Santos & Aguiar, 2019
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publication ID |
https://doi.org/ 10.5852/ejt.2019.542 |
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publication LSID |
lsid:zoobank.org:pub:3539B76F-619A-4863-997B-A449BE1BD352 |
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DOI |
https://doi.org/10.5281/zenodo.5943070 |
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persistent identifier |
https://treatment.plazi.org/id/03894A65-580D-FFD2-54AB-B35D286EF820 |
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treatment provided by |
Plazi |
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scientific name |
Mallochia aurantia ( Santos & Aguiar, 2008 ) |
| status |
comb. nov. |
Mallochia aurantia ( Santos & Aguiar, 2008) comb. nov.
Distictus aurantium Santos &Aguiar, 2008: 36–39 . Original description, figure, phylogenetic assessment.
Distictus aurantius – Yu et al. 2012 . Listed, specific epithet changed.
Comments
The considerable differences between the type species Distictus tibialis and D. aurantium Santos & Aguiar have already been stressed in Santos & Aguiar (2008: 32, 38). Those differences prompted the authors to include both species in a phylogenetic analysis based on 65 morphological characters, including 19 outgroup taxa, mostly from the group the genus was supposed to belong to, the subtribe Gabuniina ( Gabunia genus group sensu Santos 2017). The two species were recovered as sister groups and hence classified within the same genus. However, examination of further comparative material of Distictus and Mallochia shows that several species of the latter genus show character states similar to species of Distictus (see ‘Comments’ section for the genus above). The characters that consistently separate the two genera are precisely those that set D. aurantium apart from the other species of the genus, such as T1 without anterolateral tooth, fore wing crossvein 3r-m absent and vein 2Cua <0.60 × as long as crossvein 2cu-a. In fact, the species is quite similar to specimens of the Nearctic Mallochia strigosa (Cresson, 1872) . Hence, Distictus aurantium is hereby transferred to Mallochia .
Nomenclature
Yu et al. (2012) changed the neutral specific epithet aurantium to the masculine aurantius , to match Distictus , which is masculine. The ICZN (1999) states, in Article 34.2.1, that “ If a species-group name is a noun in apposition its ending need not agree in gender with the generic name with which it is combined and must not be changed to agree in gender with the generic name ”. The epithet was originally intended as a name in aposition, but this was not stated, making the emendation to aurantius acceptable. In an equivalent action, the epithet is now changed to aurantia to match Mallochia , which is feminine.
Material examined
Total of 13 specimens, 4 ♀♀ and 9 ♂♂.
Holotype
BRAZIL – Espírito Santo State • ♀; Santa Maria de Jetibá, Fazenda Clarindo Krüger ; 29 Nov.–6 Dec. 2002; MTTavares, C.O. Azevedo exped.; Malaise trap; Pt. B6; UFES.
Paratypes
BRAZIL – Espírito Santo State • 1 ♀; Fazenda Paulo Seick, Área 1 ; 3–10 Dec. 2004; M.T. Tavares exped.; Pt. B3; UFES • 1 ♂; Domingos Martins, Mata Pico do Eldorado ; 3–10 Dec. 2004; M.T. Tavares exped.; Malaise trap; Pt. T4; UFES • 1 ♂; Santa Teresa, Estação Biológica Santa Lúcia ; 9–13 May 2006; M.T. Tavares exped.; Malaise trap; Pt. B1; UFES • 2 ♂♂; Conceição do Castelo, Ribeirão do Meio ; 17–24 Mar. 2007; A.P. Aguiar exped.; near lagoon ; Malaise trap; UFES • 2 ♂♂; Domingos Martins, Mata Pico do Eldorado ; 3–10 Dec. 2004; M.T. Tavares exped.; Pt. T6; UFES .
URUGUAY • 2 ♂♂; 40 km NW of Tacuarembó; 2–9 Feb. 1963; J.K. Bouseman leg.; AMNH .
Other material
ARGENTINA • 1 ♀; Misiones, Mado, Puerto Magdalena ; 22 Oct. 1964; A. Kovac leg.; AMNH • 1 ♀; Salta, El Alisal , 45 km W of Salta; alt. 1950 m; 1–29 Dec. 1987; S. Peck and J. Peck leg.; USUC .
BRAZIL – Bahia State • 1 ♂; Ilhéus, Fazenda São José ; 9 Dec. 2003; J. Cardoso and J. Maia leg.; Malaise trap; Pt. 2; UFES .
Distribution records
Brazil (BA, ES), Argentina, Uruguay.
Keys to species of Distictus Townes, 1966 View in CoL View at ENA
Females
1. Metasoma mostly orange (RGB 226,93,5), reddish (200,52,5) or reddish-brown (175,53,8), T2–8 mostly without posterior yellow stripes ( Figs 7D View Fig , 9C View Fig , 14E View Fig ), at most T2–3 with incomplete stripe ( Fig. 6D View Fig ) .................................................................................................................................. 2
– Metasoma black (23,5,2) or dark brown (31,11,2), at least T2–3 with posterior yellow stripes ( Figs 4C View Fig , 10D View Fig , 11C View Fig ) .......................................................................................................................... 9
2. Supra-clypeal area entirely pale yellow ( Fig. 13A, C View Fig ), rarely with blackish marks ( Fig. 15 View Fig A–B); orbital band on malar space complete, rarely ventrally briefly interrupted ( Fig. 15 View Fig A–B) ............... 3
– Supra-clypeal area with blackish or reddish-brown marks ( Figs 7A View Fig , 9A View Fig ); orbital band on malar space interrupted ......................................................................................................................................... 4
3. Metapleuron entirely black ( Fig. 15C View Fig ); mesosternum entirely black; fore and mid coxae pale yellow, without dorso-apical brown spot ( Fig. 15G View Fig ); central yellow mark on mesopleurum extending to sternaulus ( Fig. 15G View Fig ); posterior area of propodeum rugulose ( Fig. 15F View Fig ); anterior margin of propodeum medially distinctly concave ( Fig. 15F View Fig ) .......................... Distictus tibialis ( Brullé, 1846) View in CoL
– Metapleuron with distict yellow mark ( Fig. 13D View Fig ); mesosternum with large yellow marks; fore and mid coxae pale yellow, usually with dorso-apical brown spot ( Fig. 13D View Fig ); central yellow mark on mesopleuron extending beyond sternaulus ( Fig. 13D View Fig ); posterior area of propodeum strigate-rugulose ( Fig. 13B View Fig ); anterior margin of propodeum medially slightly concave ( Fig. 13B View Fig ) .............................. ............................................................................................................. Distictus paratibialis sp. nov.
4. Propodeum with sublateral elongate yellow marks ( Figs 7C View Fig , 14B View Fig ); mesosoma black or reddishbrown with distinct yellow marks ( Figs 9C View Fig , 14E View Fig ); mesoscutum centrally with yellow spot; wings hyaline; epomia distinct only after diverging from pronotal collar .................................................. 5
– Propodeum without sublateral elongate yellow marks, only with propodeal crests bright yellow ( Fig. 5B View Fig ); mesosoma almost entirely black, with a few, tiny yellow spots ( Fig. 5C View Fig ); mesoscutum centrally without yellow spot; wings infuscate ( Fig. 5C View Fig ); epomia complete ..................................... ..................................................................................................................... Distictus ardens sp. nov.
5. Orbital band continuous on temple ( Fig. 9C View Fig ); mesosoma black with yellow marks; malar space blackish; scutelar carina black ( Fig. 12B View Fig ) ........................................................................................ 6
– Orbital band interrupted on temple ( Fig. 14E View Fig ); mesosoma reddish-brown with yellow marks; malar space reddish-brown; scutelar carina mostly yellow ( Fig. 14B View Fig ) ........................................................ .................................................................................................................. Distictus terrosus sp. nov.
6. Anterior transverse carina of propodeum medially broadly arched forwards ( Figs 6B View Fig , 9B View Fig ); sublateral yellow marks of propodeum comma-shaped ( Figs 6B View Fig , 9B View Fig ); T2 with posterior yellow stripe; hind femur blackish and dark brown; epicnemial carina irregular or distinctly sinuous .......................... 7
– Anterior tranverse carina of propodeum medially weakly bent ( Fig. 7C View Fig ); sublateral yellow marks of propodeum not comma-shaped ( Figs 7C View Fig , 12F View Fig ); T2 without posterior yellow stripe; hind femur mostly reddish or light orange; epicnemial carina more or less uniformly curved .......................... 8
7. Posterior area of propodeum strigate ( Fig. 6B View Fig ); small yellow marks on collar and mesopleuron ( Fig. 6D View Fig ); T3 with blackish marks and narrow posterior yellow stripe ( Fig. 6D View Fig ); hind tibia blackish, except base, orange; spiracle of T1 not prominent; clypeus black, centrally with small yellow spot ( Fig. 6A View Fig ); thyridium about as wide as long .............................................. Distictus asterios sp. nov.
– Posterior area of propodeum strigate-rugulose ( Fig. 9B View Fig ); extensive yellow marks on collar and mesopleuron ( Fig. 9C View Fig ); T3 entirely reddish ( Fig. 9C View Fig ); hind tibia deep yellow, except apical 0.1 blackish; spiracle of T1 slightly prominent; clypeus almost entirely yellow ( Fig. 9A View Fig ); thyridium distinctly wider than long ( Fig. 9B View Fig ) ...................................................... Distictus commatus sp. nov.
8. Hind femur reddish, apically blackish ( Fig. 7D View Fig ); supra-clypeal area with three blackish marks that are almost connected centrally to form an inverted Y ( Fig. 7A View Fig ); median tubercle of clypeus large; orbital band widely interrupted on malar space, broadly disconnected at eye margin ( Fig. 7A View Fig ); spiracle of T1 not prominent; propodeum sublaterally with small yellow dot basad tranverse carina ( Fig. 7C View Fig ) ....................................................................................................... Distictus ateles sp. nov.
– Hind femur fully bright orange ( Fig. 12D View Fig ); supra-clypeal area without three blackish marks that are almost connected centrally to form an inverted Y, only with two oblique blackish marks on ventral 0.3 ( Fig. 12A View Fig ); median tubercle of clypeus small; orbital band shortly interrupted on malar space, nearly fused at eye margin ( Fig. 12A View Fig ); spiracle of T1 slightly prominent; propodeum sublaterally with distinct yellow spot basad tranverse carina ( Fig. 7C View Fig ) ................................................................. ................................................................................................................. Distictus notabilis sp. nov.
9. T4–8 blackish or dark brown, with wide posterior yellow stripes ( Figs 4C View Fig , 11C View Fig ); scutellar carina yellow; hind coxa mostly yellow, dorso-apically with large dark brown mark ( Figs 4C View Fig , 11C View Fig ); sternaulus moderately deep; epicnemial carina irregular; propleuron yellow; S2–5 bright or pale yellow ............................................................................................................................................. 10
– T4–8 blackish, without posterior yellow stripes ( Fig. 10D View Fig ); scutellar carina black; hind coxa black, dorso-basally with large yellow mark ( Fig. 10D View Fig ); sternaulus deeply impressed; epicnemial carina clearly sinuous; propleuron black; S2–5 brownish, with narrow posterior pale yellow stripes ......... ..................................................................................................................... Distictus daelus sp. nov.
10. Hind tibia blackish ( Fig. 4C View Fig ); mesoscutum coriarious, matte and with lateral yellow marks; pale yellow marks on mesosoma ( Fig. 4 View Fig B–C); fore wing vein 3-M distinctly longer than 2-M; metapleuron rugulose; supra-clypeal area pale yellow, centrally with small blackish marks ( Fig. 4A View Fig ); fore and mid coxa pale yellow, with small dorso-apical brown spots; hind t1–4 whitish ................................ ................................................................................................................. Distictus apaensis sp. nov.
– Hind tibia bright yellow ( Fig. 11C View Fig ); mesoscutum densely punctate, shiny and without lateral yellow marks ( Fig. 11 View Fig B–C); bright yellow marks on mesosoma; fore wing vein 3-M about as long as 2-M; metapleuron lineate; supra-clypeal area bright yellow, without blackish marks ( Fig. 11A View Fig ); fore and mid coxa dorsally bright yellow, without dorso-apical brown spots; hind t1–4 bright yellow .......... ..................................................................... Distictus mexicanus Kasparyan & Ruíz-Cancino, 2005 View in CoL
Males
1. Metasoma mostly orange (RGB 226,93,5) ( Fig. 15E View Fig ), reddish (200,52,5) or reddish-brown (175,53,8) ( Fig. 14D View Fig ), rarely one or more of T2–4 with posterior yellow stripe ( Figs 8D View Fig , 10F View Fig ); hind tibia entirely yellow, or blackish only at apex .......................................................................................... 2
– Metasoma blackish, all tergites with wide posterior yellow stripe (as in Fig. 11C View Fig ); hind tibia deep yellow with basal 0.1 blackish .................... Distictus mexicanus Kasparyan & Ruíz-Cancino, 2005 View in CoL
2. Orbital band continuous on temple ( Fig. 15E View Fig ); scutellar carina black; mesosoma black or dark brown with extensive yellow marks ( Fig. 10F View Fig ) ........................................................................................... 3
– Orbital band interrupted on temple ( Fig. 14D View Fig ); scutellar carina almost entirely yellow; mesosoma reddish-brown with extensive yellow marks ( Fig. 14D View Fig ) .................................................................... .................................................................................................................. Distictus terrosus sp. nov.
3. T2–3 without posterior yellow stripes, rarely T2 with posterior yellow stripe ................................ 4
– T2–3 with posterior yellow stripes ................................................................................................... 6
4. Metapleuron rugulose; propodeum posteriorly rugulose ou strigate-rugulose (as in Fig. 15F View Fig ); propleuron entirely pale yellow; mid coxae entirely pale yellow; T1 mostly orange, sometimes blackish; malar space pale yellow .................................................................................................... 5
– Metapleuron lineate; propodeum posteriorly distinctly striate (as in Fig. 7C View Fig ); propleuron entirely black, sometimes with small posterior yellow spot; mid coxae ventrally blackish; T1 mostly black; malar space black ......................................................................................... Distictus ateles sp. nov.
5. Metapleuron fully black ( Fig. 15E View Fig ), without yellow mark; mesosternum entirely black, rarely with small pale yellow marks; scape dorsally brown, sometimes apically with small yellow spot ........... ..................................................................................................................... Distictus tibialis View in CoL sp. nov.
– Metapleuron black with distinct yellow mark (similar to Fig. 13D View Fig ); mesosternum almost or entirely pale yellow; scape dorsally pale yellow, rarely light brown ............................................................... ............................................................................................................. Distictus paratibialis sp. nov.
6. Anterior transverse carina of propodeum almost straight ( Fig. 8B View Fig ); T4 with posterior pale yellow stripe ( Fig. 8 View Fig C–D); pronotum polished smooth along collar ( Fig. 8C View Fig ); propleuron mostly yellow; sternaulus moderately deep ( Fig. 8C View Fig ); mesopleuron shiny and weakly striate on dorsal area between hypoepimeron and epicnemial carina ..................................................... Distictus caligaris sp. nov.
– Anterior transverse carina of propodeum centrally distinctly arched forwards ( Fig. 10G View Fig ); T4 without posterior yellow stripe ( Fig. 10F View Fig ); pronotum distinctly striate along collar; propleuron entirely dark brown; sternaulus deeply impressed (as in Fig. 10D View Fig ); mesopleuron matte and almost entirely lineate .......................................................................................................... Distictus daelus sp. nov.
| UFES |
Brazil, Universidade Federal do Espirito Santo, Departamento de Biologia, Colecao Entomologica |
| AMNH |
USA, New York, New York, American Museum of Natural History |
| USUC |
USA, Colorado, Fort Collins, Colorado State University |
| UFES |
Universidade Federal do Espirito Santo |
| AMNH |
American Museum of Natural History |
No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.
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Mallochia aurantia ( Santos & Aguiar, 2008 )
| Supeleto, Fernanda A., Santos, Bernardo F. & Aguiar, Alexandre P. 2019 |
Distictus aurantium Santos &Aguiar, 2008: 36–39
| Distictus aurantium Santos &Aguiar, 2008: 36–39 |
Distictus aurantius –
| Yu et al. 2012 |