Cleistepyris, Kieffer, 1910

Marques Jr, Wilson J., Azevedo, Celso O. & Colombo, Wesley D., 2025, Revision of the Brazilian Northeastern Cleistepyris (Hymenoptera: Bethylidae), with description of six new species, Zootaxa 5692 (1), pp. 1-30 : 3

publication ID	https://doi.org/10.11646/zootaxa.5692.1.1
publication LSID	lsid:zoobank.org:pub:E48DDDA4-1E5B-4D6C-900A-785F513FD064
DOI	https://doi.org/10.5281/zenodo.17362699
persistent identifier	https://treatment.plazi.org/id/0388E840-5835-1D72-FF13-FE961AB9F94C
treatment provided by	Plazi
scientific name	Cleistepyris
status

Treatment

Key for males from Northeastern Brazil

1. Mandible with one distinct ventralmost tooth, remaining teeth fused............................................ 2

- Mandible with all teeth distinct.......................................................................... 8

2. Median clypeal lobe trapezoidal ( Fig. 6B View FIGURE 6 ).................................................................. 3

- Median clypeal lobe angulate ( Fig. 6D View FIGURE 6 )................................................................... 5

3. Frons and dorsal pronotal area coriaceous; apical harpal margin slightly incurved ( Figs 11I, J View FIGURE 11 ); cuspis ventral arm setose ( Figs 11A, B View FIGURE 11 ).................................................................. C. santacatarinae ( Evans, 1964)

- Frons and dorsal pronotal area smooth; apical harpal margin with small, well-delimited excavation ( Figs 11A, B View FIGURE 11 ); cuspis ventral arm glabrous........................................................................................ 4

4. Aedeagal inner lobe as wide and longer than outer lobe; outer lobe pointed................. C. quadratus ( Evans, 1963)

- Aedeagal inner lobe twice narrower, twice shorter than outer lobe; outer lobe rounded or slightly concave........................................................................................... C. paraensis ( Kieffer, 1910b)

5. Metapostnotal median carina complete ( Fig. 8A View FIGURE 8 ); apical harpal margin strongly emarginated to half of harpe ( Figs 11G, H View FIGURE 11 )....................................................................................... C. piripiri sp. nov.

- Metapostnotal median carina incomplete posteriorly ( Fig. 8F View FIGURE 8 ); apical harpal margin not strongly emarginate to half of harpe. ................................................................................................... 6

6. Emargination of apical harpal margin less than 0.15× harpe length ( Fig. 11A, B View FIGURE 11 )................... C. awaguaja sp. nov.

- Emargination of apical harpal margin more than 0.25× harpe length............................................. 7

7. Dorsal harpal margin outcurved; outer lobe apex rounded and posterior to inner lobe apex.......................................................................................... C. quelatus ( Ramos, Araújo & Azevedo, 2010)

- Dorsal harpal margin straight; outer lobe apex acute and anterior to inner lobe apex................ C. tremembe sp. nov.

8. Median clypeal lobe angulate ( Fig. 6B View FIGURE 6 ).................................................................... 9

- Median clypeal lobe trapezoidal ( Fig. 6D View FIGURE 6 )................................................................. 13

9. Mandible teeth with less than five distinct apical teeth; frons and dorsal pronotal area coriaceous; apical harpal margin outcurved.................................................................... C. neotropicus ( Evans, 1963)

- Mandible teeth with five distinct apical teeth; frons and dorsal pronotal area smooth; apical harpal margin slightly incurved or strongly emarginate.................................................................................. 10

10. Harpe with medial-dorsal corner partially arched, apical margin strongly emarginate; cuspis with dorsal arm laminar; cuspis with ventral arm glabrous (see Fig. 18 in Azevedo & Batista 2002)............ C. megaventris ( Azevedo & Batista, 2002)

- Harpe whole arched, bilaminar, apical margin slightly incurved; cuspis with dorsal arm bilaminar; cuspis with ventral arm setose ( Figs 11E, F View FIGURE 11 ).................................................................................. 11

11. Anterior ocellus anterior to supra-ocular line ( Fig. 5A View FIGURE 5 ); posterior hypopygeal margin straight ( Fig. 10E View FIGURE 10 )..................................................................................... C. ocullatus ( Azevedo & Batista, 2002)

- Anterior ocellus posterior to supra-ocular line ( Fig. 5B View FIGURE 5 ); posterior hypopygeal margin incurved ( Fig. 10F View FIGURE 10 ).............. 12

12. Aedeagal apical lobe acute apically, dorsal surface slightly convex; aedeagus with two pairs of apical lobes (see figs 5–7 in Leal & Azevedo 2001)........................................................ C. clypeatus ( Leal & Azevedo, 2001)

- Aedeagal apical lobe narrowing apicad, dorsal surface strongly convex; aedeagus with one pair of apical lobe ( Figs 11E, F View FIGURE 11 )...................................................................................... C. kanela sp. nov.

13. Frons and dorsal pronotal area coriaceous; digitus glabrous; dorsal harpal margin straight.......................... 14

- Frons and dorsal pronotal area smooth; digitus densely setose; dorsal harpal margin outcurved....................... 15

14. Cuspis with ventral arm rounded ( Fig. 11F View FIGURE 11 and see fig. 77 in Evans 1963)....................... C. inca ( Evans, 1963)

- Cuspis with ventral arm straight ( Fig. 11J View FIGURE 11 )................................................ C. tenetehara sp. nov.

15. Metapectal-propodeal disc with transverse posterior carina, metapostnotal median carina complete; aedeagus with two pairs of apical lobes..................................................................... C. spinipes ( Evans, 1969)

- Metapectal-propodeal disc without transverse posterior carina, metapostnotal median carina incomplete; aedeagus with one pair of apical lobes................................................................... C. guajajara sp. nov.