Kateretidae, Erichson, 1846

Hisamatsu, Sadatomo, 2011, A review of the Japanese Kateretidae fauna (Coleoptera: Cucujoidea), Acta Entomologica Musei Nationalis Pragae 51 (2), pp. 551-585 : 555

Key to Kateretidae genera of Japan

1 Tarsal claws not dentate or slightly dilated at base ( Figs. 3J , 5E , 9E ). Apical segment of maxillary palpus slender ( Figs. 3I , 5F , 8G , 9F ). Ovipositor with styli ( Figs. 3D , 4G , 5D , 6F , 7G , 8D , 9D ). ............................................................................................................. 2

– Tarsal claws distinctly dentate at base ( Figs. 10F , 11H ).Apical segment of maxillary palpus enlarged ( Figs. 10E , 11G ). Ovipositor without styli ( Figs. 10G , 11D ). ......................... 5

2 Abdominal sternite III or III–V with tuft of setae in the middle in both sexes ( Figs. 5H , 9K ). Ovipositor with short styli ( Figs. 5D , 9D ). ............................................................ 3

– Abdominal sternite III or III–V without tuft of setae. Ovipositor with long styli ( Figs. 3D , 4G , 6F , 7G , 8D ). ............................................................................................................. 4

3 Color reddish-yellow to dark brown. Lateral margins of pronotum smooth. Male protibia strongly arcuate ( Figs. 2D , 9I ). Abdominal sternite III with tuft of setae in middle ( Fig. 9K ). ...................................................................................... Sibirhelus Kirejtshuk, 1989

– Color black. Lateral margins of pronotum distinctly serrate ( Fig. 5G ). Male protibia straight. Abdominal sternites III–V with tuft of setae in middle ( Fig. 5H ). ............................ ................................................................................................. Platamartus Reitter, 1892

4 Pronotum rounded at posterior angles ( Fig. 3H ). Dorsal pronotal disc surface with coarse, large punctures, larger than eye-facet. Abdominal tergite VIII in males concealed. ........ ...................................................................................................... Kateretes Herbst, 1793

– Pronotum angulate at posterior angles ( Figs. 6H–N , 7I , 8F ). Dorsal pronotal disc surface with dense, small punctures, smaller than eye-facet. Abdominal tergite VIII in males exposed. .................................................................... Heterhelus Jacquelin du Val, 1858

5 Dorsal pronotal disc surface with sparse punctures, and sparsely covered with short setae ( Fig. 2E ). Pronotum narrower than elytra at base; posterior angles obtuse ( Fig. 10H ); hind margin straight. ................................................................ Brachypterus Kugelann, 1794

– Dorsal pronotal disc surface with dense punctures, and densely covered with long setae ( Fig. 2F ). Pronotum as wide as elytra at base; posterior angles angulate ( Fig. 11F ); hind margin sinuate. ............................................................ Brachypterolus Grouvelle, 1913

Figures

Fig. 3. Kateretes japonicus Hisamatsu, 1985.A – tegmen (ventral view); B – median lobe (ventral view); C – median lobe (lateral view); D – ovipositor; E – right protibia; F – right antenna of female (segments 1–3); G – right antenna of male; H – pronotum; I – left maxillary palpus; J – tarsal claw (right foreleg). A–J = Nagano Pref., Tokugo Pass.

Fig. 5. Platamartus jakowlewi Reitter, 1892.A – tegmen (ventral view); B – median lobe (ventral view); C – median lobe (lateral view); D – ovipositor; E – tarsal claw of right foreleg; F – left maxillary palpus; G – pronotum; H – abdominal sternites; I – right antenna, female; J – ditto, male. A–J – Tochigi Pref., Watarase-yûsuichi.

Fig. 9. Sibirhelus corpulentus (Reitter, 1900).A – tegmen (ventral view); B – median lobe (ventral view); C – median lobe (lateral view); D – ovipositor; E – tarsal claw of right foreleg; F – left maxillary palpus; G – male right antenna; H – female right protibia; I – male right protibia; J – pronotum; K – abdominal sternites. A–K = Tochigi Pref., Watarase-yûsuichi,

Fig. 8. Heterhelus (Boreades) solani (Heer, 1841). A – tegmen (ventral view); B – median lobe (ventral view); C – median lobe (lateral view); D – ovipositor; E – right antenna, male; F – pronotum; G – left maxillary palpus.A–G = Hokkaido, Kawayu.

Fig. 4. Kateretes takagii S-T. Hisamatsu, 2006. A – tegmen (ventral view); B – median lobe (ventral view); C – median lobe (lateral view). D–E – antennal segments 1–2 (lateral view): D – male, E – female. F – right protibia; G – ovipositor.A–G = Hokkaido, Piasiri moore.

Fig. 6. Heterhelus (Heterhelus) scutellaris (Heer, 1841).A – tegmen (ventral view); B – median lobe (ventral view); C – median lobe (lateral view); D – parameres of the lectotype of H. japonicus; E – median lobe of the lectotype of H. japonicus (lateral view); F – ovipositor; G – male abdominal sternite VII; H–N – variation of pronotum (H – Hokkaido; I – Aomori; J – Niigata; K – Tokyo; L – Yamaguchi; M – Ehime; N – Fukuoka); O – male right antenna. A, B, C, G, O = Hokkaido, Hitsujigaoka; F = Ehime Pref., Omogokei.

Fig. 7. Heterhelus (Heterhelus) morio (Reitter, 1878). A – tegmen (ventral view); B – median lobe (ventral view); C – median lobe (lateral view); D – male right antenna; E – parameres of the lectotype of H. morio; F – median lobe of the lectotype of H. morio (lateral view); G – ovipositor; H – male abdominal sternite VII; I – pronotum. A, B, C, D, G, H, I = Ehime Pref., Omogokei.

Fig. 10. Brachypterus urticae (Fabricius, 1792). A – tegmen (ventral view); B – median lobe (ventral view); C – median lobe (lateral view); D – male right antenna; E – left maxillary palpus; F – tarsal claw of right foreleg; G – ovipositor; H – pronotum.

Fig. 11. Brachypterolus pulicarius (Linnaeus, 1758). A – tegmen (ventral view); B – median lobe (ventral view); C – median lobe (lateral view); D – ovipositor; E – male right antenna; F – pronotum; G – left maxillary palpus; H – tarsal claw of right foreleg. A–H = Aomori Pref., Azumahama.

Fig. 2. Habitus and labels of Kateretidae spp. A–B – Heterhelus (Heterhelus) morio (Reitter, 1878): A – lectotype, male; B – ditto, labels; C – Heterhelus (Boreades) solani (Heer, 1841), male (Hokkaido); D – Sibirhelus corpulentus (Reitter, 1900), male (Iwate Pref.); E – Brachypterus urticae (Fabricius, 1792), male (Hokkaido); F–G – Brachypterolus shimoyamai Hisamatsu, 1985: F – holotype, male; G – ditto, labels. Scale bars = 1.0 mm.