Occidenchthonius

Zaragoza, Juan A., 2017, Revision of the Ephippiochthonius complex in the Iberian Peninsula, Balearic Islands and Macaronesia, with proposed changes to the status of the Chthonius subgenera (Pseudoscorpiones, Chthoniidae), Zootaxa 4246 (1), pp. 1-221 : 128-129

Key to adults of the Occidenchthonius species

1 Chelal hand chaetotactic formula 4:4:4, seta ih5 absent. Cheliceral hand lyrifissure ldb present ( O. machadoi -group)...... 2

- Chelal hand chaetotactic formula 4:5:4, seta ih5 present. Cheliceral hand lyrifissure ldb present or absent.............. 11

2 Epigean species. Well developed eyes, anterior and posterior eyes with convex lens................................ 3

- Hypogean species. Eyeless or posterior eyes reduced to eye-spots.............................................. 4

3 Fixed chelal finger with 12–16 teeth and movable with 7. Chela stouter: (♂) 4.5, (♀) 3.8–4.7 times longer than deep, length.............................................................................. O. machadoi ( Vachon, 1940)

- Fixed chelal finger with 20–21 teeth, movable 9–10. Chela more slender: (♂) 5.1, (♀) 4.6–4.9 times longer than deep.................................................................................... O. canariensis ( Beier, 1965)

4 Carapace with a total of 2‒8 preocular and ocular microsetae.................................................. 5

- Carapace with a total of 16 preocular and ocular microsetae............................... O. setosus ( Mahnert, 1993)

5 Posterior margin of carapace with 2 (exceptionally 3) setae................................................... 6

- Posterior margin of carapace with 4 (very exceptionally 3 or 5) setae.......................................... 10

6 Eyes absent.............................................................................. O. oromii n. sp.

- Eyes present, at least anterior eyes with weak lens, posterior eyes reduced to indistinct spots......................... 7

7 Dorsal face of chelal hand without lyrifissure hd .................................. O. gracilimanus ( Mahnert, 1997)

- Dorsal face of chelal hand with lyrifissure hd .............................................................. 8

8 Distal half of movable chelal finger with 9–10 pointed teeth up to slightly proximad of trichobothrium st, basal half with 8–12 rounded and fused vestigial teeth up to level slightly proximad of trichobothria sb ................................. 9

- Distal half of movable chelal finger with 12–15 pointed teeth up to halfway between trichobothria st and sb, basal half with 10–12 rounded and fused vestigial teeth up to halfway between trichobothria sb and b ................. O. mahnerti n. sp.

9 Anterior medial margin of carapace prominent; cheliceral spinneret absent in male; stouter and smaller pedipalp: femur (♂) 6.0–6.1, (♀) 5.9 times longer than broad, length (♂) 0.53 mm, (♀) 0.53–0.58 mm; hand (♂) 2.2–2.3, (♀) 2.1 times longer than deep; chela (♂) 5.7–5.8, (♀) 5.0–5.2 times longer than deep, length (♂) 0.71–0.72 mm, (♀) 0.75‒0.76 mm .. O. beieri n. sp.

- Anterior medial margin of carapace nearly straight; cheliceral spinneret present in male, hump like; pedipalp femur (♂) 6.8– 7.2, (♀) 6.0–6.9 times longer than broad, length (♂) 0.62–0.65 mm, (♀) 0.63–0.72 mm, hand (♂) 2.4–2.6, (♀) 2.2–2.5 times longer than deep, chela (♂) 5.9–6.5, (♀) 5.5–6.2 times longer than deep, length (♂) 0.79–0.88 mm, (♀) 0.84–0.95 mm .............................................................................. O. tamaran (Mahnert, 2011)

10 Smaller, chela length (♀) 0.69 mm, 5.0–5.5 times longer than deep; trichobothrium ist slightly distad to esb .............................................................................................. O. lopezi (Mahnert, 2011)

- Larger, chela length (♀) 1.08 mm, 6.1 times longer than deep; trichobothrium ist distinctly distad of esb .............................................................................................. O. tenerifae (Mahnert, 2011)

11 Movable cheliceral finger with isolated subapical tooth (di).................................................. 12

- Movable cheliceral finger without isolated subapical tooth (di)............................................... 24

12 Posterior margin of carapace with 4 macrosetae............................................................ 13

- Posterior margin of carapace with 2 macrosetae........................................................... 15

13 Epigean species with normal eyes; carapace with 2 preocular microsetae on each side; fixed chelal finger with 12–16 triangular teeth (not including proximal rounded teeth); smaller: chela length (♂) 0.45–0.57 mm, (♀) 0.50–0.66 mm ............. 14

- Endogean or hypogean species, eyes reduced or absent; carapace with 1 preocular microseta on each side; fixed chelal finger with 17–21 triangular teeth (not including proximal rounded teeth); larger: chela length (♂) 0.89–1.12 mm, (♀) 0.84–1.37 mm ................................................................................ O. cassolai (Beier, 1973)

14 Proximal half of fixed chelal finger with 2–5 rounded teeth merging into an evident high bulge of marginal lamina........................................................................................ O. parmensis ( Beier, 1963)

- Proximal half of fixed chelal finger with 2–5 rounded and spaced teeth, marginal lamina weakly raised................................................................................................ O. berninii (Callaini, 1983)

15 Without eyes or eye-spots............................................................................ 16

- With eyes, anterior pair with convex lens, posterior pair reduced to spots....................................... 23

16 Trichobothrium ist distinctly distad of esb ................................................................ 17

- Trichobothrium ist proximad of esb ....... O. morenoi ( Carabajal Márquez, García Carrillo & Rodríguez Fernández, 2011)

17 Trichobothrium ist proximad of lyrifissure fb ............................................................. 18

- Trichobothrium ist distinctly distad of lyrifissure fb ........................................................ 20

18 Larger and more slender: (♀) pedipalpal femur 7.5 times longer than broad, length 0.88 mm; chela (♀) 7.3 times longer than deep, length 1.23 mm ............................................................ O. sendrai (Zaragoza, 1985)

- Smaller and less slender: (♀) pedipalpal femur 5.8–6.5 times longer than broad, length 0.52–0.62 mm; chela (♀) 5.2–5.5 times longer than deep, length 0.70–0.86 mm .................................................................. 19

19 Chelicera with 6 setae and 4–5 lateral microsetae on hand................................. O. thaleri ( Gardini, 2009)

- Chelicera with 6 setae and only 1 lateral microseta on hand............................... O. minutus ( Vachon, 1940)

20 Tergite I with 2 setae................. O. ambrosiae ( Carabajal Márquez, García Carrillo & Rodríguez Fernández, 2012)

- Tergite I with 4 setae................................................................................ 21

21 Cheliceral hand with 6 setae (excluding microsetae)........................................................ 22

- Cheliceral hand with 7–8 setae (excluding microsetae)........................................................................................... O. bullonorum ( Carabajal Márquez, García Carrillo & Rodríguez Fernández, 2012)

22 Larger and slender: pedipalpal femur (♀) 7.5–7.7 times longer than broad, length (♀) 0.93–1.00 mm; chela (♀) 5.9–6.0 times

longer than deep, length (♀) 1.27–1.37 mm ............................................... O. felgueraorum n. sp. - Shorter and less slender: pedipalpal femur (♀) 5.5 times longer than broad, length 0.56 mm; chela (♀) 5.2 times longer than deep, length 0.78 mm ................................................. O. villacarrillo ( Zaragoza & Pérez, 2013)

23 Lyrifissure ldb present on cheliceral hand, close to seta db. Trichobothrium ist slightly distad of esb and well proximad of lyri- fissure fb ................................................................................. O. anae n. sp.

- Lyrifissure ldb absent on cheliceral hand. Trichobothrium ist well distad of esb and slightly proximad of lyrifissure fb ..................................................................................... O. dubius ( Mahnert, 1993)

24 Posterior row of carapace with 2 macrosetae.............................................................. 25

- Posterior row of carapace with 4 macrosetae......................................... O. cardosoi (Zaragoza, 2012)

25 Chelal hand lyrifissure hp absent....................................................................... 26

- Chelal hand lyrifissure hp present...................................................................... 27

26 Carapace without preocular microsetae.................................................... ( O. verai -group) 28

- Carapace with 2 preocular microsetae on each side....................................... O. pinai (Zaragoza, 1985)

27 Smaller, epigean, species: pedipalp femur (♀) 5.1–5.3 times longer than broad, length 0.44–0.47 mm; chela (♀) 3.7–3.9 times longer than deep, length 0.61 mm .......................................................... O. serranoi n. sp.

- Larger, hypogean, species: pedipalp femur (♀) 8.7–8.8 times longer than broad, length 1.28–1.30; chela (♀) 7.5–7.9 times lon- ger than deep, length 1.72–1.82............................................................ O. ortunoi n. sp.

28 Subterranean species without eyes...................................................................... 29

- Epigean species with eyes, anterior pair with convex lens........................................ O. gardinii n. sp.

29 Chelicera with 6 macrosetae and 1–2 (rarely 3) lateral microsetae. Trichobothrium ist distinctly distad of esb ........... 30

- Chelicera with 6 macrosetae and 4–5 lateral microsetae. Trichobothrium ist level with esb ................................................................................................ O. giennensis ( Zaragoza & Pérez, 2013)

30 Smaller. Chela length about 0.70 mm or less.............................................................. 31

- Larger. Chela length about 1.00 mm or more.............................................................. 32

31 Fixed chelal finger with 14–15 teeth; stouter and shorter pedipalp: femur (♂) length 0.39–0.42 mm, 4.8–5.0 times longer than broad, chela (♂) length 0.58–0.60 mm, 4.8–5.0 times longer than deep........................... O. montagudi n. sp.

- Fixed chelal finger with 19 teeth; more slender and longer pedipalp: femur (♂) length 0.54 mm, 6.0 times longer than broad, chela (♂) length 0.71 mm, 5.7 times longer than deep.......................................... O. lencinai n. sp.

32 Trichobothrium ist proximad of or level with lyrifissure fb; smaller species, pedipalpal femur (♀) 5.7–7.5 times longer than broad, length 0.66–0.94 mm, chela (♀) length 0.94–1.29 mm ................................................ 33

- Trichobothrium ist distinctly distad of lyrifissure fb; larger species, pedipalpal femur (♀) 7.9–8.6 times longer than broad, length about 1.10 mm, chela (♀) length 1.40–1.53 mm ......................................................41

33 Movable chelal finger with 3–8 rounded vestigial teeth without dental canals in basal half......................... 34

- Movable chelal finger with 9–12 rounded vestigial teeth without dental canals in basal half........................ 37

34 Trichobothrium ist distinctly proximad of lyrifissure fb ..................................................... 35

- Trichobothrium ist level with lyrifissure fb ..................................................... O. riopar n. sp.

35 Lyrifissure ldb present on cheliceral hand. O. cazorlensis Carabajal Márquez, García Carrillo & Rodríguez Fernández, 2001

- Lyrifissure ldb absent on cheliceral hand................................................................. 36

36 More slender species, pedipalpal femur (♀) 6.9–7.1 times longer than broad......................... O. mateui n. sp.

- Stouter species, pedipalpal femur (♀) 6.1 times longer than broad................................ O. hoerwegi n. sp.

37 Cheliceral fixed finger with 2 large distal teeth............................................................ 38

- Cheliceral fixed finger with only 1 large distal tooth........................................................................................... O. torremarinae ( Carabajal Márquez, García Carrillo & Rodríguez Fernández, 2012)

38 Smaller: (♀) pedipalpal femur length 0.66–0.75 mm, 5.7–6.4 times longer than broad; chela (♀) length 0.94–1.10 mm, 5.2–5.5 times longer than deep.................... O. perezi ( Carabajal Márquez, García Carrillo & Rodríguez Fernández, 2011)

- Larger: (♀) pedipalpal femur length 0.80–0.99 mm, 6.4–7.5 times longer than broad; chela (♀) length 1.06–1.38 mm, 5.7–6.5 times longer than deep............................................................................... 39

39 Male spinneret prominent........................................................... O. verai (Zaragoza, 1985)

- Male spinneret absent or strongly reduced................................................................ 40

40 Lyrifissure ldb of cheliceral hand present.............................................. O. ventalloi ( Beier, 1939)

- Lyrifissure ldb of cheliceral hand absent.................................... O. espanyoli ( Zaragoza & Pérez, 2013)

41 Movable chelal finger with 14 vestigial teeth without dental canals in basal half....................... O. murcia n. sp.

- Movable chelal finger with only 6–8 vestigial teeth without dental canals in basal half............................ 42

42 Lyrifissure ldb of cheliceral hand present................................................................................................... O. ruizporteroae (Carabajal Márquez, García Carrillo & Rodríguez Fernández, 2001) .

- Lyrifissure ldb of cheliceral hand absent......................................................... O. ebo n. sp.