Andreaea clavata Broth., Bibliotheca Botanica

Suárez, Guillermo M., 2024, Taxonomical status of Andreaea clavata var. tenuior Broth. (Bryophyta, Andreaeaceae), Phytotaxa 642 (3), pp. 225-228 : 225

Andreaea clavata Broth., Bibliotheca Botanica 87: 10. 1916.

Type:— Bolivia. Felsen am Huaillattanisee, 4900 m, T. Herzog 2958 (lectotype, designated here: JE 04000662!). Bolivia. Cochabamba: an Felsen im Schneetälchen des Cerro Tunari, 5100 m, T. Herzog 4785 (paratypes: JE 04000657!, PC 0703449 [digital image!], available at http://colhelper.mnhn.fr/, W 1914-0004183 [digital image!], available at https://www.jacq.org/#database).

= Andreaea clavata var. tenuior Broth., Bibliotheca Botanica 87: 10. 1916. Type:— Bolivia. Cochabamba: Felsen am Huaillattanisee, 4900 m, T. Herzog 2960 (lectotype, designated here: JE 04000659!, isolectotypes: PC 0703448 [digital image!], available at http:// colhelper.mnhn.fr/, W 1914-0004137 [digital image!] available at https://www.jacq.org/#database), syn. nov.

Observations: the name on the original label of the specimen is “ Andreaea perimbricata Broth. ”

Note— Andreaea clavata is characterized by the large, red-brown, and clavate plants with stem not or occasionally branched, in transverse section rounded, rather homogeneous, and central strand absent. The axillary hairs are 2-celled, brownish. The leaves are crowded and imbricate when dry, erect-patent when wet, straight, obovate, more rarely panduriform, very concave, cochleariform, with obtuse to rounded, straight apex, cucullate, erose. The margin is entire and the costa is absent. Lamina is unistratose, with upper cells ± isodiametric, rounded to subquadrate, rarely shortly oblong, with thin walls with prominent hyaline cuticular dorsal papillae. The mid cells are shortly oblong or rectangular and the basal cells short to long rectangular. ( Fig. 1 , 2 ).

Figures

FIGURE 1. Andreaea clavata. A. Habit of plant in wet; B. Habit of plant in dry; C. leaves; D. Tip of leaf; E. Marginal median leaf cells; F. Basal leaf cells; G. Leaf cross section; H. Axillary hairs; I. Cross section of stem. (All from Lectotype, Herzog 2958, JE).

FIGURE 2. Detailed pictures of Andreaea clavata. A. Leaf; B. Leaf apex showing upper leaf cells; C. Mid leaf cells; D. Basal leaf cells. (All from Lectotype, Herzog 2960, JE).