Medina, Anichtchenko & Sciaky & Medina, 2022

Anichtchenko, Alexander, Sciaky, Riccardo & Medina, Milton Norman, 2022, Taxonomic notes on the genus Hexagonia Kirby, 1825 with description of three new species from the Philippines (Coleoptera: Carabidae: Hexagoniini), Baltic Journal of Coleopterology 22 (1), pp. 55-66 : 62-63

Medina sp. n. ( Fig. 4 )

Type material: Holotype, female – “ Philippines, Mindanao , Gingoog | Eureka Mt., 730m, 8°41.4970N | 125°14.2710E, 4.v.2022 | M. Medina leg.” ( DUBC).

Diagnosis. Easily distinguishable from all known species in the Oriental region in the combination of metallic blue elytra and strongly trapezoid head with the fovea behind the eye connected by a shallow groove with the postorbital pore.

Description. Body length: 11 mm. Width/length of head:1.13; width/length of pronotum:1.1; length/ width of elytra: 1.7. Head and pronotum black with weak bluish sheen, elytra black with bright blue luster. Mandibles dark brown, palpi brown, antenna and legs uniformly black. Lower surface black.

Head strongly trapezoid and depressed, slightly narrower than the pronotum. Eye moderately large and markedly protruded, 0.5 times as long as temples; temples protruding; posterior border almost perpendicular; constriction of neck weak and shallow, neck narrow. Mandibles elongate, almost straight in basal two thirds, incurved towards apex. Palpi elongate, narrowed towards apex, smooth. Mentum with wide triangular tooth, bisetose. Antenna short, attaining the lateral pore of pronotum; three basal antennomeres and basal half of antennomere 4 glabrous. Clypeal suture shallow, laterally indistinct; clypeus slightly emarginate at apex. Labrum with deep wide triangular incision, asymmetric, right lobe longer. Frons with two irregularly shaped, shallow impressions. Supraorbital pore connected by a shallow groove with the postorbital pore. Surface with extremely fine transverse microreticulation, iridescent.

Pronotum wider than the head, obovate, widest at apical quarter; dorsal surface depressed.Anterior margin slightly convex, anterior angles completely rounded. Lateral margins evenly curved, immediately in front of base slightly excised; base straight, basal angles obtuse and slightly produced. Median line deep, impunctate, almost attaining anterior margin and base; both transverse sulci obsoleted. Basal grooves starts right from the posterior angles in form of short, smooth, diagonal impressions. Anterior margin and base not margined, lateral borders narrowly margined, marginal sulcus slightly wider near lateral pore. Anterior lateral seta situated at widest point, posterior lateral seta absent. Dorsal surface smooth, with some very shallow irregular transverse sulci, with extremely fine transverse microreticulation, slightly iridescent. Elytra elongate, markedly depressed, parallel sided. Base oblique, humeri rounded; lateral margins straight. Basal margin straight and forming a curve at the junction with lateral margin, between striae 4 and 5. Scutellar stria long and connected with stria 1. All striae deeply impressed and moderately punctate. Intervals flat and smooth. Scutellar seta situated at origin of stria 1. Interval 3 with three setiferous pores, the anterior pore situated in the stria 3 at the level of the end of scutellar stria; the discal pore situated in apical third of elytra near stria 2; posterior one situated close to apex, near stria 2. Interval 5 of elytra with one pore located opposite the middle between discal and apical pores. Marginal series consisting of 12–13 pores. Intervals impunctate and with weak microreticulation, consisting of polygonal meshes, very glossy.

Legs rather short, tarsomere 3 of all tarsi two times shorter than basal one; tarsomere 4 wide and very deeply excised, with dense brush­like pilosity on ventral surface. Tarsal claws edentate. Three basal tarsomeres of all legs triangular, very wide and dorsally smooth.

Male unknown.

Female genitalia. Gonocoxite 1 apically with five elongate setae along ventro­apical rim. Gonocoxite 2 rather narrow, triangular, slightly curved, with obtusely rounded apex; with three short and stout ensiform setae on the external­apical edge and one large suboval setiferous pit with seta near apex on the medio­ventral surface.

Distribution. Known only from the type locality, situated in the province of Misamis Oriental. in the north of the island of Mindanao.

Etymology. This new species is named after ‘ Panumanod ’, the guardian of the animals of the Higaunon Tribe in Northern Mindanao.

Affinities. This species is very peculiar within the genus in the metallic color and the strongly trapezoid head, with the supraorbital pore connected by a groove with the postorbital pore. In these two characters it is strongly reminiscent of some species of the Madagascan genus Omphreoides Fairmaire, 1896 ( Jeannel, 1948). The following species ( H. nigrocyanea n. sp.) is clearly similar in size, shape and general characters, but some of the features are less evident (the trapezoid head, the metallic color, ecc.). These species seem to form a small natural group within the genus, apparently without relationships with any other species group.

Figures

Fig. 4. Habitus of Hexagonia panumanod sp. n. Holotype, female. Scale bar 5 mm.