Amphidraus zipa, Galvis, William, 2017

Galvis, William, 2017, Nineteen new species of Amphidraus Simon, 1900 (Salticidae: Euophryini) from Colombia, with comments about their conservation, Zootaxa 4286 (1), pp. 1-40 : 26-29

Amphidraus zipa sp. nov.

Figs 15a–d , 19c–d , 23f–g , 25f, 25l , 27

Types. Holotype: male from Camino Real , Parque Nacional Natural Tamá, Santander, Colombia, 2500 m [7.297492°N, 72.263947°W], VI.1999, E. González ( ICN –Ar 3363) . Paratype: one female from Sendero Binacional , Parque Nacional Natural Tamá, Santander, Colombia, 2470 m [7.297492°N, 72.263947°W], unknown coll. ( ICN –Ar 3364).

Etymology. The epithet is a noun in apposition that honors one of the four supreme rulers of the Muisca Confederation, ancient inhabitants of the central Andean highlands of the present-day Colombia’s Eastern Range: the Zipa , the Zaque , the Iraca and the Tundama .

Diagnosis. Males of A. zipa sp. nov. are similar to those of A. tundama sp. nov by the shape of the RTA, the presence of basal widening of the embolus (bwe), and the absence of a dorsal tibial spur (DTS) in the palp. However, they can be distinguished from those by the curved and longer embolus, the presence of a bilobed process on the embolic disc (PED), the presence of a proventral and proximal femoral widening (PvFw, black arrow in Figs 15a , 23g ), and a pointed basal widening of the embolus (bwe) ( Figs 15a–b , 23f–g ). Females of A. zipa sp. nov. are similar to those of A. tundama sp. nov. by the general disposition of the internal structures of the epigyne, but can be recognized for the their longer copulatory ducts, larger spermathecae, and the absence of the bilobed medial and posterior pocket in the epigyne ( Figs 15c–d , 25f, 25l ).

Description. Male (holotype). Total length: 2.96. Carapace brown, 1.47 long, 0.72 wide, 0.92 high ( Fig. 19c ). OC black, 0.83 long. Anterior eye row 1.18 wide and posterior 1.02 wide. Sternum brown, 0.56 long, 0.52 wide. Labium brown, 0.16 long, 0.23 wide. Chelicerae brown with five retromarginal and two promarginal teeth. Palp brown with long white hairs on patella and tibia, with slender and curved RTA, a proventral and proximal femoral widening (PvFw), presence of basal widening of the embolus, bilobed PED and a straight TL ( Figs 15a–b ). Legs 4312, all yellow with dark brown marks. Leg macrosetae: femur, I d 1 di; II d 1 di, p 1 di; III d 2 di, p 1 di; IV d 2 di; patella, III–IV r 1 me; tibia, I v 2-2 -2; II v 1-2 -2; III v 1 -0-1, p 1-0-1, r 1-0-1; IV v 1 -0-1, p 0-1-1, r 0-1-1; metatarsus, I–II v 2-2; III v 2 -0-1, p 1-0-1, r 2 di; IV v 1 di, p 1-0-2, r 1-1-2. Abdomen brown with a reticulated dark brown pattern of marks, and medial transversal light brown stripes ( Fig. 19c ).

Female (paratype). Total length 3.78. Carapace dark brown, 1.71 long, 1.24 wide, 0.78 high ( Fig. 19d ). OC black, 0.81 long. Anterior eye row 1.16 wide and posterior 1.09 wide. Sternum yellow, 0.67 long, 0.54 wide. Labium yellow, 0.11 long, 0.23 wide. Chelicerae yellowish, with four retromarginal and two promarginal teeth.

Legs 4312, all yellow. Leg macrosetae: femur, II–IV d 1 di; patellae, III r 1 me; tibiae, I v 2-2 -2; II v 2-2 -2, p 1 di; III–IV v 1 pr, p 1-1, r 1-1; metatarsus, I v 2-2; II v 2-2, p 1 di; III–IV v 1 -0-1, p 1-0-2, r 1-0-2. Abdomen gray with a dark brown reticulated pattern of marks, and transversal lighter stripes ( Fig. 19d ). Epigyne ( Figs 15c–d , 25f, 25l ) with anterior copulatory openings, a posterior single medial pocket, long copulatory ducts and rounded spermathecae.

Distribution. Colombia (Santander) ( Fig. 27 ). Known altitudinal distribution: 2470–2500 m.

Figures

FIGURES 15 a – d. A. zipa sp. nov., a) left male palp, retrolateral view (black arrow showing the proventral femoral and proximal widening of the palpal femur, PvFw); b) same, ventral view; c) epigyne, cleared, ventral view; d) same, dorsal view. Abbreviations: bwe = basal widening of the embolus, cd = copulatory duct, co = copulatory opening, e = embolus, ED = embolic disc, EF = embolic filament, fd = fertilization duct, PED = process on embolic disc, po = epigynal pocket, RTA = retrolateral tibial apophysis of the male palp, SD = sperm duct, sp = spermatheca, TL = tegular lobe. Scale bars: 0.30 mm (a – b), 0.20 mm (c – d).

FIGURES 19 a – d. A. zaque sp. nov., a) male holotype (ICN – Ar 3378); b) female paratype (ICN – Ar 3379). A. zipa sp. nov., c) male holotype (ICN – Ar 3363); d) female paratype (ICN – Ar 3364). Scale bars: 0.50 mm.

FIGURES 23 a – g. Amphidraus spp, left male palp. A. tundama sp. nov., a) ventral view; b) same, retrolateral view; c) palpal femur, showing the proventral bump (PvFB, black arrow), ventral view. A. zaque sp. nov., d) ventral view; e) same, retrolateral view. A. zipa sp. nov., f) ventral view; g) same, showing the proventral femoral widening (PvFw, black arrow) in the femur, retrolateral view.

FIGURES 25 a – l. Amphidraus spp, epigyne. A. sikuani sp. nov., a) ventral view; g) disposition of the internal structures. A. somondoco sp. nov., b) ventral view; h) disposition of the internal structures. A. sua sp. nov., c) ventral view; i) disposition of the internal structures. A. tundama sp. nov., d) ventral view; j) disposition of the internal structures. A. zaque sp. nov., e) ventral view; k) disposition of the internal structures. A. zipa sp. nov., f) ventral view; l) disposition of the internal structures.

FIGURE 27. Distribution records of the species of the genus Amphidraus Simon from Colombia treated here. Symbols with a dot inside represent species that have distributions in the Cundiboyacense high-Andean plateau, in the Boyacá and Cundinamarca departments.