Blepharoneura thetis Hendel, 1914

Norrbom, Allen L. & Condon, Marty, 2010, Revision of the femoralis group of Blepharoneura Loew (Diptera: Tephritidae) 2374, Zootaxa 2374 (1), pp. 1-139 : 113-115

Blepharoneura thetis Hendel

Figs. 67–68, 94, 198–199

Blepharoneura thetis Hendel 1914: 21 [in key], 22 [description]; Aczél 1950: 198 [in catalog]; Foote 1967: 18 [in catalog]; Norrbom et al. 1999: 106 [in catalog].

Diagnosis. This species can be distinguished from other species of Blepharoneura by the following combination of characters: Anepisternum entirely yellow; scutum with pair of brown submedial vittae, extended to trapezoidal brown mark on posterior margin, but without sublateral vittae; scutellum with band or pair of transverse brown marks extending to side and to basal seta but not to basal margin; cell r1 proximally with single relatively narrow, tapering marginal hyaline mark; cell r2+3 with 2 marginal hyaline marks, posterior one not connected to marginal hyaline spot in cell r4+5; cell r4+5 with 2 small marginal hyaline spots or one bilobed mark (spots connected proximally). It is very similar to B. fernandezi and a probably undescribed species (see sp. nr. thetis ), which differ in lacking brown scutellar markings and having only 1 marginal hyaline spot in cell r4+5. The female is unknown.

Description. Head: Dark brown area on ocellar tubercle extended less than half distance to postocellar seta. Large brown spot surrounding medial vertical seta extended ventromesally to level of postocellar seta [may connect to form band, medial part not visible in Nova Teutonia male, head is detached and glued to mounting card]. Occipital suture narrowly orange brown.

Thorax ( Fig. 94 ): Scutum nonmicrotrichose except posterior to dorsocentral seta and laterally, with pair of broad dark brown submedial presutural vittae, broadest slightly anterior to level of posterior margin of postpronotal lobe, extended to posterior brown mark. Posterior margin with broad elongate trapezoidal dark brown mark, extended laterally to intra-alar seta and lateral to base of scutellum. Scutellum with complete transverse band (Nova Teutonia ♂) or pair of irregular, transverse, lateral brown marks (holotype) including basal marginal seta but not extending to basal margin, continuing (at least in Nova Teutonia ♂) on side to ventral margin (but ventrobasal corner yellow). Subscutellum yellow. Mediotergite with pair of broad, narrowly separated red brown vittae. Pleuron entirely yellow. Basalare entirely yellow. Dorsocentral seta aligned slightly posterior to postalar seta.

Legs: Entirely yellow.

Wing (Figs. 67–68): Length 5.80–5.94 mm, width 2.90–3.07 mm, ratio 1.94–2.00. Crossvein r-m at 0.55– 0.57 distance from bm-cu to dm-cu. Cell c with 2 rectangular hyaline spots, both reaching costa and subcosta; medial brown area slightly paler than area of cell r1 posterior to pterostigma, sometimes fainter posteriorly, and approximately as broad as to slightly narrower than hyaline spots. Pterostigma with large subapical hyaline spot [#3] reaching R1. Cells r1 and r2+3 basally (proximal to apex of R1) without hyaline spots except for minute spot in r1 posterior to apex of vein Sc. Radial cells medially with tapering basal marginal hyaline mark in cell r1 [#5] and aligned spot in r2+3 [#8] forming acute triangular to nipple-shaped mark, extending at least to R4+5 and sometimes into r4+5, but spot in cell r4+5 [#14] sometimes slightly more distal; spot in r4+5 [#14] touching R4+5 but extended less than halfway across cell; cell r2+3 with more distal moderately large hyaline spot [#9], touching R4+5 and R2+3; cell r4+5 with medial hyaline spot [#15] near dm-cu small. Distally cell r1 with 1 marginal subapical hyaline spot [#6]. Cell r2+3 with 2 elongate marginal hyaline marks [#10, 11], both extending to vein R4+5. Cell r4+5 with small hyaline spot [#16] on anterior margin aligned between marginal marks in cell r2+3, and with 2 small marginal hyaline spots [#18, 18A], on right wing of Nova Teutonia male connected on proximal ends. Cell m with 3 small hyaline spots, 2 marginal [#27, 29] and 1 anteromedial [#26 and/or #26A], none extended to vein M. Cell br with subbasal hyaline spot [#12]. Cell bm with small circular subbasal and larger circular subapical spots [#19, #20]. Posteromedial part of wing with hyaline spots mostly isolated; cell br with subapical spot [#13] and usually (except 1 wing of Nova Teutonia ♂) with smaller more proximal posterior spot [#44]; cell dm with subbasal hyaline band [fused #51, #52] aligned or partially aligned with more proximal spot in cell br [#44] and proximal anterior spot [#31] in cell cu1 (when latter spots present) sometimes (Nova Teutonia ♂) narrowly connected anteriorly or posteriorly to more distal spots in dm; cell dm also usually (except 1 wing of Nova Teutonia ♂) with anterior medial spot [#21] aligned with subapical spot [#13] in cell br, and with broad posteromedial spot [fused #22, #23, #24]; cell cu1 medially sometimes (holotype) with 3 anterior hyaline spots [#31, #32, #33] and 2 marginal spots, medial anterior spot [#32] elongate and almost connected to marginal spot, or (Nova Teutonia ♂) with or without proximal anterior spot [#31] and with 2 distal anterior spots [#32, #33] fused with medial marginal spot [#36] to form Y-shaped mark aligned with posteromedial spot in cell dm; proximal marginal spot [fused #34, #39] across apex of vein A1+Cu2 isolated; medial marginal mark (or posterior part of Y-shaped mark) partially divided and relatively broad [fused #36, #36A?] (holotype) or narrower and undivided [#36 only] (Nova Teutonia ♂); subapical marginal hyaline mark [#37] small. Cell dm with subapical hyaline spot [#25] moderate sized, not extended to vein M.

Abdomen: All tergites with pair of broad dark brown markings forming irregular vittae; mark on each tergite extended broadly along posterior margin to lateral margin of tergite; spots broadly separated medially by tapering, nearly straight margined yellow area; brown spots solid except for 1 small anterior yellow spot in spots on tergites 3–5.

Male terminalia: Medial surstylus with prensisetae subequal, separated by width or slightly more than width of medial prensiseta, medial prensiseta on short lobe ( Figs. 198–199 ).

Distribution. Brazil (Santa Catarina, Rio Grande do Sul).

Type data. Holotype ♂ ( NMW), BRAZIL: Rio Grande do Sul, [no date], Stieglmayer [examined]. The holotype was not marked as such, but it has a label with " Blepharoneura thetis H." in Hendel's writing. It is a holotype by monotypy, as Hendel stated that he examined just one male specimen. We added a holotype label.

Other specimens examined. BRAZIL: Santa Catarina: Nova Teutonia , 27°11'S 52°23'W, 2 Jan 1953, F. Plaumann, 1♂ ( BMNH USNMENT00213840 ) .

Figures

FIGURES 94–97. Thorax, dorsal: 94, B. thetis (holotype); 95, B. bivittata (Costa Rica: S Upala, USNMENT00054202); 96–97, B. brevivittata (Costa Rica: Rincon de Osa, USNMENT00054203; Costa Rica: Golfo Dulce, USNMENT00048518).

FIGURES 196–199. Epandrium and surstyli, lateral and posterior: 196–197, B. femoralis (Brazil: Nova Teutonia); 198– 199, B. thetis (holotype).