Amplirhagada alicunda, Köhler, 2011

Köhler, Frank, 2011, Descriptions of New Species of the Diverse and Endemic Land Snail Amplirhagada Iredale, 1933 from Rainforest Patches across the Kimberley, Western Australia (Pulmonata: Camaenidae), Records of the Australian Museum 63 (2), pp. 167-202 : 173-175

Amplirhagada alicunda n.sp.

Type locality. Western Australia, NW Kimberley, 27 km SE Walsh Point , ca. 16 km SSW of Mt. Connor, 13 km from coast; 14°41'45"S 126°04'04"E ( RFS 11-4 , coll. V. Kessner, 11 Jun 1987) ( Fig. 1 ) .

Type material. Holotype WAM S34710 (preserved specimen) (Pl. 1.3; Table 1) . Paratypes WAM S34711 (2 preserved specimens) , FMNH 220384 (2 preserved specimens) , AM C.472921 (10 dried shells), WAM S34712 (16 dried shells) , FMNH 220385 (25 dried shells) .

Etymology. Species epithet derived from “alicunde” (Latin = from somewhere, from any place), in reference to the remote type locality, far away from any named place.

Description

Shell (Pl. 1.3; Fig. 8 A–D ). Broadly conical to semi-globose with elevated spire. Solid, not translucent. Periphery slightly compressed to slightly angulate; upper sector of whorl flattened, basal sectors rounded. Umbilicus 70–100% concealed by columellar reflection. Background colour crème to horn; peripheral band conspicuous, usually well developed, rather thin, clearly visible on most whorls; sub-sutural band broad, diffuse to well developed, brown, clearly visible on most whorls; outer and inner lip colour white, conspicuously contrasting shell colour. Protoconch 2.4 mm in diameter, comprising about one whorl, with very weak radially elongated pustulations. Teleoconch with faint, regular axial growth lines, evenly distributed across shell. Angle of aperture about 60 degrees; outer lip moderately thick, well rounded, slightly expanded, not reflected; basal node absent or weak. Parietal wall of inner lip very inconspicuous. Average shell size 16.0±1.4 × 20.2± 0.7 mm ( Table 1).

Radular and jaw morphology ( Fig. 8 E–G ). Tooth formula C + 16–18 + 4 + 18–20 (n = 1). Jaw with 12 plates.

Genital morphology ( Figs 9–10 ). Penis straight, more or less of same length as anterior part of oviduct; distal part of penis proper narrow, proximal part inflated. Penial sheath delicate. Length of penial retractor muscle equivalent to about ¼ of length of penial complex. Penial verge extending about 1/10 of length of penial chamber, slender with pointed tip. Proximal part of inner penial wall entirely covered by extremely fine pustulation. Main stimulatory pilaster strongly enlarged, comprising proximal half of inner penial chamber, cone-shaped, sculptured by regularly spaced, lateral ridges that support dense rows of little hooks. Vas deferens entering penial sheath within proximal third of penial complex.Vagina moderately long, tubular; inner vaginal wall and inner wall of bursa copulatrix with continuous, well-developed, smooth longitudinal pilasters. Bursa copulatrix short, reaching base of spermoviduct. Free oviduct rather straight comprising less than half of length of anterior part of oviduct. Spermoviduct longer than anterior part of oviduct.

Aestivation strategy and ecology. Free sealer, inhabits locally restricted vine thicket.

Remarks. Anatomical description based on dissection of one specimen. Listed by Solem (1991) as “ Amplirhagada NSP 19”. Shell similar to other relatively large, broadly conical species with spiral bands, such as A. combeana Iredale, 1938 and A. mitchelliana Solem, 1981 but anatomy of inner penial wall clearly different.

Figures

Figure 1. Distribution of Amplirhagada species described in the present work. (A) A. atlantis n.sp. (B) A. carsoniana n.sp. (C) A. alicunda n.sp. (D) A. davidsoniana n.sp. (E) A. vialae n.sp. and A. discoidea n.sp. (F) A. forrestiana n.sp. (G) A. inusitata n.sp. and A. epiphallica n.sp. (H) A. lindsayae n.sp. (I) A. angustocauda n.sp. (K) A. moraniana n.sp. (L) A. gardneriana n.sp. (M) A. bendraytoni n.sp. (N) A. coffea n.sp. (P) A. carinata Solem, 1981. When species occur at more than one locality, the type locality of newly described taxa is marked with an asterisk.

Figure 8. Amplirhagada alicunda n.sp.SEM of shell (AM C.472921), radula and jaw (WAM S34711). (A) Protoconch and first teleoconch whorls viewed from above. (B) Detail of first teleoconch whorls viewed from above. (C) First teleoconch whorls viewed obliquely from above. (D) Close-up showing sculpture on last whorl. (E) Jaw. (F) Central and inner lateral teeth viewed from above. (G) Marginal teeth viewed from above. Scale bars: A–D, 1 mm; E, 100 µm; F, 10 µm, G, 20 µm.

Figure 9. Genitalia of Amplirhagada alicunda n.sp.(WAM S34711). For labelling of structures see Fig. 3. Scale bar = 5 mm.

Figure 10. Penial anatomy of Amplirhagada alicunda n.sp. (WAM S34711). For labelling of structures see Fig. 4. Scale bar = 3 mm.