Amplirhagada epiphallica, Köhler, 2011

Köhler, Frank, 2011, Descriptions of New Species of the Diverse and Endemic Land Snail Amplirhagada Iredale, 1933 from Rainforest Patches across the Kimberley, Western Australia (Pulmonata: Camaenidae), Records of the Australian Museum 63 (2), pp. 167-202 : 193-195

publication ID	https://doi.org/ 10.3853/j.0067-1975.63.2011.1581
publication LSID	lsid:zoobank.org:pub:1BCD4085-D2B9-400D-B504-8C85C30303D6
persistent identifier	https://treatment.plazi.org/id/0384503A-FFA1-3C42-FC06-FBCAFE90699A
treatment provided by	Felipe
scientific name	Amplirhagada epiphallica
status	sp. nov.

Treatment

Amplirhagada epiphallica View in CoL n.sp.

Type locality. Western Australia, NW Kimberley, N side of Prince Frederick Harbour , near mouth, 16 km W of Mt. Anderdon, 0.3 km from coast; 14°57'10"S 125°16'30"E ( RFS 14-1 ; coll. V. Kessner, 14 Jun 1987) ( Fig. 1 View Figure 1 ) GoogleMaps .

Type material. Holotype WAM S34745 View Materials (preserved specimen) (Pl. 1.15; Table 1) . Paratypes WAM S34746 View Materials (3 preserved specimens) , FMNH 200539 About FMNH (4 preserved specimens) , WAM S34747 View Materials (2 dried shells) , FMNH 220541 About FMNH (2 dried shells) .

Etymology. The species epithet refers to the presence of an epiphallus, which is a very distinctive feature among species of Amplirhagada .

Description

Shell (Pl. 1.15; Fig. 43 A–C View Figure 43 ). Broadly conical with low spire, almost discoid. Thin to delicate, translucent. Periphery rounded to angulate; upper and basal sectors of whorls well rounded. Umbilicus 10–50% concealed by columellar reflection. Background colour pale yellowish brown; peripheral band well marked, moderately thick, brown, visible on most whorls; sub-sutural well marked to diffuse, moderately thick, visible on most whorls; outer and inner lip colour whitish. Protoconch 2.5 mm in diameter, comprising about 1.2 whorls, essentially smooth. Teleoconch with regular axial growth lines; evenly distributed across shell surface. Angle of aperture about 45 degrees; outer lip thin, well rounded, expanded, not reflected; basal node absent or weakly developed. Parietal wall of inner lip inconspicuous. Average shell size 13.7±0.8 × 20.1± 0.7 mm ( Table 1).

Radular and jaw morphology ( Fig. 43 D–F View Figure 43 ). Tooth formula C + 13–14 + 4 + 18–20, with 150 rows of teeth 130 (n = 1). Jaw with 11 plates.

Genital morphology ( Figs 44–45 View Figure 44 View Figure 45 ). Penis complex elongated, tubular, coiled; actual penis extending proximal two thirds of penial complex. Inner penial wall completely lined with regular, fine pustulation arranged in corrugated pilasters; no main stimulatory pilaster differentiated. Penial sheath thick. Penial retractor muscle stubby, no longer than 1/10 of penial complex. Vas deferens moderately thick, entering penial sheath at apical end of penial complex, forming an extended loop, reflecting as an elongate, thin, tubular epiphallus, which gives rise to the short penis. Inner wall of epiphallus smooth, ciliated. Penial verge tiny, slender, forming a loop with pointed tip. Vas deferens elongated, winding before entering the uterus. Vagina long, tubular; inner vaginal wall and wall of bursa copulatrix with smooth longitudinal pilasters. Bursa copulatrix slightly extending base of spermoviduct, with bulbous head. Free oviduct elongated, coiled before entering the uterus, comprising more than half of length of anterior part of oviduct. Spermoviduct no longer than anterior part of oviduct. Albumen gland as long as spermoviduct.

Aestivation strategy. Free sealer.

Remarks. Anatomical description based on dissection of the holotype. Listed by Solem (1991) as “ Amplirhagada NSP 35” and found in sympatry with A. inusitata n.sp. and A. pusilla Solem, 1981 . The flat shell is similar to that of A. discoidea . However, the genital anatomy of this species is very unusual for Amplirhagada showing a combination of exclusive features, such as a very long and winding vas deferens, a long tubular vagina, very long tubular penis and vas deferens reflecting as an epiphallus within the strongly developed penial sheath, and a short, thin and coiled penial verge.