Ademosyne bacca Ponomarenko, 1969

Yan, Evgeny V., Beutel, Rolf G. & Ponomarenko, Alexander G., 2017, Ademosynidae (Insecta: Coleoptera): A new concept for a coleopteran key taxon and its phylogenetic affinities to the extant suborders, Palaeontologia Electronica 5 (8), pp. 1-22 : 12

Ademosyne bacca Ponomarenko, 1969 A. bacca : Ponomarenko, 1969, p. 133

Figure 3.5, 3.6 , Figure 4.4, 4.5

Material. Holotype: PIN 2240 /310, beetle without antennae and legs; Eastern Asia , Dzhailyaucho, Lower Triassic . Paratype: PIN 2240 /294 from the same locality .

Revised description. Pronotum very short, posterior margin weakly bisinuated. Mesocoxae twice as wide as long, two times broader than space between them. Metaventrite strongly transverse, half as wide as maximum length at posterior margin, which is twice as wide as anterior margin; paracoxal suture straight, mesal portions of metacoxae weakly shifted posteriorly. Fifth abdominal ventrite 1.5 times longer than penultimate. Ventral side of body covered with small punctures. See measurements in Table 1.

Differs from other species in the shape of the pronotum, which is twice as wide as long; presence of very deep transverse procoxal rests ( Figure 4.4 , cx.r) of mesoventrite; metaventral paracoxal suture strongly shifted towards posterior margin.

Figures

FIGURE 3. Genera of Ademosynidae family, habitual photos. 1, Gnathosyne akkolkensis Ponomarenko, 1969, holotype, PIN 2496/7 counterpart. 2, Ademosyne kirgizica Ponomarenko, 1969, holotype PIN 2096/1369. 3, 4, A. kirgizica Ponomarenko, 1969 paratype PIN 2555/1696 print and counterprint. 5, A. bacca Ponomarenko, 1969, holotype PIN 2240/310. 6, A. bacca Ponomarenko, 1969, paratype PIN 2240/294. 7, A. ellyptica Ponomarenko, 1969, holotype PIN 2240/224. 8, Cephalosyne capitata Ponomarenko, 1969 holotype PIN 2240/302. Scale bars equal 1 mm.

FIGURE 4. Genera of Ademosynidae family, line drawings. 1, Gnathosyne akkolkensis Ponomarenko, 1969, holotype, PIN 2496/7 counterpart. 2, Ademosyne kirgizica Ponomarenko, 1969, holotype PIN 2096/1369. 3 A. kirgizica Ponomarenko, 1969, paratype PIN 2555/1696 print and counterprint. 4, A. bacca Ponomarenko, 1969 holotype PIN 2240/310. 5, A. bacca Ponomarenko, 1969, paratype PIN 2240/294. 6, A. ellyptica Ponomarenko, 1969, holotype PIN 2240/224. 7, Cephalosyne capitata Ponomarenko, 1969, holotype PIN 2240/302. Scale bars: 1 mm. Label abbreviations: c.e – compound eye; crd – cardo; cx.r – coxal rests; cx1, 2 – pro- and mesocoxae; ep – epipleura; epm2 – mesepimeron; fm1 – profemur; gul – gular plate (gula); i.p1 – prosternal intercoxal process; l.d – pronotal longitudinal depression; l.i – lateral incixions of submentum; l.r – longitudinal ridge; md – mandibles; msc – mesoscutum; mt – mentum; ms.p – mesosternal pit; o.f – occipital foramen; par – parietal; pd – pedicellus; pl1 – propleurite; p.mt – prementum; p.t.p. – posterior tentorial pit; sc – scapus; scl – scutellum; sct – mesoscutum; s.g – gular suture; s.l2 – longitudinal suture of mesoventrite; s.l3 – longitudinal suture of metaventrite; smt – submentum; s.pc – paracoxal suture.

FIGURE 5. Petrosyne Ponomarenko, 1969 and ademosynid beetles re-assigned to Polyphaga suborder, photos and line drawings. 1, 2 Petrosyne liassica Ponomarenko, 1969, holotype PIN 166/39. 3, 4 Ranis collevus Ponomarenko, 1969, holotype PIN 371/1755 and line drawing. 5, 6, R. ovalis Ponomarenko, 1969, holotype PIN 2066/2642. Scale bars equal 1 mm. Label abbreviations: cx.r – coxal rests; msc – mesoscutum; tb1 – protibia.

FIGURE 6. Comparative morphology of †Ademosynidae, polyphagan beetles and stem group Coleoptera. 1, 2, Stenocyphon sasaji Lawrence, 2001. 3, 4, Epiphanis cornutus (Eschscholtz, 1829). 5, 6, Dolichosyne sulcata Ponomarenko, 1969. 7, 8 Gnathosyne akkolkensis Ponomarenko, 1969. 9, 10, Archosyne permiana Ponomarenko et al., 2014. 11, 12, Tecticupes heckeri Rohdendorf, 1961 (†Taldycupedidae). 13, 14, Permocupes sojanensis Ponomarenko, 1963 (†Permocupedidae). Scale bars equal 1mm. Label abbreviations: mb.c – membranous connection.