Orchis markusii Tineo, Pl. Rar. Sicil.

Barone, Giulio, Domina, Gianniantonio, Gristina, Emilio Di, Mirabile, Giulia & Gargano, Maria Letizia, 2023, Typification of names published by Vincenzo Tineo from Sicily, Phytotaxa 612 (2), pp. 139-147 : 142-143

7. Orchis markusii Tineo, Pl. Rar. Sicil. [Ed. 2] 1: 9(–10). 1846. [15 March 1846]

≡ Orchis pseudosambucina Ten. subsp. markusii (Tineo) Nyman, Consp. Fl. Eur. 4: 693. 1882. [October 1882] ≡ Orchis romana Sebast. f. markusii (Tineo) Soó, Repert. Spec. Nov. Regni Veg. 24(1–3): 30. 1927. [25 May 1927] ≡ Orchis sulphurea Sims var. markusii (Tineo) Jahand. & Maire, Cat. Pl. Maroc 1: 151. 1931. [after 1 August 1931] ≡ Dactylorhiza romana (Sebast.) Soó var. markusii (Tineo) Soó, Ann. Univ. Sci. Budapest. Rolando Eotvos, Sect. Biol. 3: 337. 1960. [November 1960], comb. inval. ≡ Dactylorhiza romana (Sebast.) Soó var. markusii (Tineo) Soó , Nom. Nov. Gen. Dactylorhiza: 3. 1962. ≡ Dactylorhiza markusii (Tineo) H.Baumann & Künkele, Mitteilungsbl. Arbeitskreis Heimische Orchid. Baden-Württemberg 13(4): 461. 1981. [28 December 1981] ≡ Dactylorhiza romana (Sebast.) Soó subsp. markusii (Tineo) Holub, Folia Geobot. Phytotax. 19(2): 214. 1984. [31 May 1984]

Ind. Loc.:—“In sylvatis montosis herbosis. Palermo a Gibilrossa, nel boschetto dè sugheri presso il Convento”.

Lectotype (here designated):—[Sicily] Orchis makusii Tin, Sopra Gibilrossa nel Boschetto dè Sugheri , 1 Aprile 1835, s. c. [manu V. Tineo] ( PAL80920 !, image available at: http://147.163.8.207/zoomify/view_img.asp?ic=80920).

Note:— Pedersen (2006: 425) reports “Type: Italy, Sicily, Palermo, Gibilrossa, 1827, Tineo s.n. (holotype?PAL, isotype FI (photo seen)”. However, no holotype was cited by Tineo (1846: 9–10) in the protologue (see Art. 9.6 of ICN); moreover, several herbarium samples from different collections prior to the publication of the name were found (preserved in FI and PAL). FI002724 bears two specimens by Vincenzo Tineo and the following labels: “ Orchis markusii Tin. Gibilrossa Boschetto di Querce, da Tineo Luglio 1848 ” and “ Orchis markusii Tin, Palermo a Gibilrossa Tin., da Tineo in maggio 1847 ”. These specimens were checked by L. Hautzinger in February 1974, who added a label with the indication “ Lectotype ”; no nomenclatural acts, however, were published. In our opinion, we cannot be certain that FI specimens are original material of the name because the date of collection is missing and the dates of arrival in FI are later than 1846. Three specimens are housed at PAL and they were collected by V. Tineo in Gibilrossa in 1834 (PAL87510) and in 1835 (PAL87511, PAL80920). We here designate the specimen PAL80920 as lectotype of the name Orchis markusii . It includes one individual, complete in all its part, and bears a label in Tineo’s handwriting. PAL80920 matches the protologue and corresponds to the current application of the name, which refers to a taxon accepted by some authors ( Wucherpfennig 2004, Pacifico & Biagioli 2016). Morphologically, it is considered distinct from Dactylorhiza romana ( Sebastiani 1813: 12) Soó (1962: 3) (the basionym, Orchis romana Sebast. , was lectotypified by Iamonico et al. 2020) or regarded as its synonym by other authors (Bullini et al. 2001, Pedersen 2006, Delforge 2016) based on molecular data.